Ann Cusack ( ; born May 22, 1961) is an American actress. She had roles in Multiplicity (1996), A League of Their Own (1992), and The Informant! (2009). Additionally, she has made guest appearances in a number of television series, including Grey's Anatomy, Scandal, One Tree Hill, Charmed, Ghost Whisperer, The Unit, Boston Legal, Bones, Frasier, Ally McBeal, Star Trek: Enterprise, Criminal Minds, Private Practice, Fargo, Better Call Saul, The Boys, and The Good Doctor.

Early life Cusack was born in New York City and was raised in Evanston, Illinois. She has four siblings, including Joan and John. Her mother, Ann Paula "Nancy" (née Carolan), was a mathematics teacher and political activist. Her father, Dick Cusack, was an actor, producer, and writer. With her sister Joan and brother John, she trained at the Piven Theatre WorkshopPiven Theatre Workshop - Home in Evanston, and graduated from the Berklee College of Music in Boston, Massachusetts, in 1987.

Career Acting Cusack starred in her first film role in 1992, when she was cast in A League of Their Own, as Shirley Baker. She was in the 1996 comedy Multiplicity, which starred Michael Keaton and Andie MacDowell. Also in 1996, she replaced Anita Barone on The Jeff Foxworthy Show until the series ended. In 1998, Cusack starred in Maggie on Lifetime Television.

Cusack has made guest appearances in Grey's Anatomy, Scandal, One Tree Hill, Charmed, Ghost Whisperer, The Unit, Boston Legal, Bones, Frasier, Ally McBeal, Criminal Minds, Private Practice, and Castle among others. In 2002, she guest-starred in the episode of Star Trek: Enterprise entitled "Carbon Creek". She had a small role in Grosse Pointe Blank, which starred her siblings John and Joan; appeared in the movie Accepted, in which she plays Diane Gaines, mother to the lead character played by Justin Long; and appeared in the film What Planet Are You From?. She also co-starred in Ace Ventura Jr: Pet Detective as Melissa Robinson Ventura, the titular character's mother (replacing Courteney Cox from the original film).

She also appeared in the SciFi series Lost Room as Helen Ruber (two episodes, 2006). In 2015, she appeared in the first episode of Fargo'''s second season. In 2016, she played Donna Dent in the Clint Eastwood-directed Warner Bros. film Sully, about Chesley Sullenberger and the events of Flight 1549, starring Tom Hanks. She also made her first appearance as Rebecca Bois, Chuck's ex wife, in season two of Better Call Saul'', later returning in season three.

Filmography

Film Year Title Role Notes1992Shirley Baker1993MaliceWaitress1994Renaissance ManBill's Secretary1995Tank GirlSub Girl1995The Point of Betrayal1996TV Woman in Van1996MultiplicityNoreen1996My Fellow AmericansWhite House Tour Guide1997Grosse Pointe BlankAmy1997Cannes ManKitty Monaco1997Peoria BabylonCandy Dineen1997Midnight in the Garden of Good and EvilDelivery Woman1999StigmataDr. Reston2000What Planet Are You From?Liz2001America's SweetheartsAssistant to Lee Phillips2006AcceptedDiane Gaines2006Deanna2006ArcAngela Blake2007Cake: A Wedding StoryCeleste2007Jenny2009Robin Mann2009The First TimeBeatrice2010Crooked LaneAva2014NightcrawlerLinda2016SullyDonna Dent2017Heart, BabyNurse Claire

Television Year Title Role Notes1992OverexposedMarcy LevinTV film1992StephanieTV series1993Love & WarKatherineEpisodes: "Valentine's Day" & "Croton on Hudson"1993Victim of Love: The Shannon Mohr StoryTelevision film1995Murder, She WroteMargaret BarkleyEpisode: "The Scent of Murder"1996–1997Karen FoxworthyMain role1998From the Earth to the MoonJan ArmstrongTV miniseries1998–1999MaggieMaggie DayLead role1999Ladies ManDelilahEpisode: "Thanks for Nothing"2000Melissa SchifferEpisode: "Kidnapped"2001Ally McBealRebecca MooreEpisode: "The Obstacle Course"2001What About Joan?Ann GallagherEpisode: "Joan's Sister Visits"2001Black RiverMandy PruellTV film2002My Sister's KeeperGraceTV film2002Family LawNancy EmersonEpisode: "Alienation of Affection"2002Star Trek: EnterpriseMaggieEpisode: "Carbon Creek"2003MiraclesKaren LongviewEpisode: "The Friendly Skies"2003FrasierAntoniaEpisode: "Daphne Does Dinner"2003Julie ShawMain role2004One Tree HillDr. Nora ThorpeEpisodes: "The Living Years" & "Hanging by a Moment"2004, 2005CharmedMiss DonovanEpisodes: "Charmed Noir" & "Carpe Demon"2005Six Feet UnderLinda HoviakEpisode: "Dancing for Me"2006Ghost WhispererGrace DowlingEpisode: "Melinda's First Ghost"2006Grey's AnatomyAmyEpisode: "What Have I Done to Deserve This?"2006Fatal Contact: Bird Flu in AmericaDenise ConnellyTelevision film2006BonesBrianna's LawyerEpisode: "The Titan on the Track"2006Close to HomeMartha HarrisEpisode: "There's Something About Martha"2006Helen RuberTV miniseries2007Boston LegalDr. Donna FolletteEpisode: "Angel of Death"2007Army WivesHannah WhiteEpisode: "Dirty Laundry"2008Brothers & SistersJamieEpisode: "Double Negative"2008GreekKaren CartwrightEpisode: "47 Hours and 11 Minutes"2008–2009Susan GilliumRecurring role2009Ace Ventura Jr.: Pet DetectiveMelissa VenturaTelevision film2009–2011Private PracticeSusan GrantRecurring role2010Criminal MindsSarah HillridgeEpisode: "Mosley Lane"2010TraumaEpisode: "Scope of Practice"2010In Plain SightMelissa DonaldsonEpisode: "Whistle Stop"2010Jane3 episodes2011Harry's LawLynette ZalesEpisode: "The Fragile Beast"2012Body of ProofGail WhirleyEpisode: "Identity"2012Hart of DixieAnnie HattenbargerEpisodes: "The Race & the Relationship" & "Disaster Drills & Departures"2012ScandalCorinne StarkEpisode: "Hunting Season"2013Masters of SexHarrietEpisode: "Brave New World"2014Sullivan & SonEllenEpisode: "Everybody Loved Frank"2015BackstromSandy Hale-CooperEpisode: "Love is a Rose and You Better Not Pick It"2015FargoJudge MundtEpisode: "Waiting for Dutch"2015, 2016CastleRitaEpisodes: "XX" & "Dead Red"2016GrimmMrs. BaskeEpisode: "Lycanthropia"2016–2018Better Call SaulRebecca Bois5 episodes2017TycoonCatherine BlakeTV series2017Mr. MercedesOlivia Trelawney 3 episodes2018Castle RockWarden Theresa Porter5 episodes2019–2026The BoysDonna JanuaryRecurring role2020Station 19JoanEpisode: "I Know This Bar" 2021 The Good DoctorIlana ReevesEpisode: "Expired" 2024 RipleyEmily Greenleaf Episode: "A Hard Man to Find"

Personal life Cusack married actor Jim Piddock on April 1, 2023 in a private ceremony in their home in Los Angeles, California. Cusack is the stepmother to Piddock's daughter from his previous marriage.

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