The Kotel'nich Tetrapod Assemblage is a tetrapod assemblage zone (also called a biozone) in Russia. Its fossils are from "a red-coloured predominantly clay-argillite-siltstone sequence exposed on the right bank of the Vyatka River south of the town of Kotel’nich". It is dated to the late Capitanian age of the Middle Permian. The type locality (Kotel'nich, or Kotel'nich 1) is in the Vanyushonki Member of the Vyatka Formation. The upstream Port Kotel'nich (Kotel'nich 2) locality, dominated by dicynodonts of genus Australobarbarus, was traditionally also included in this assemblage; however, this is now questioned due to fossil and possible time differences.
Paleobiota Listed from Suchkova et al. (2026) unless otherwise cited.
Synapsids
Therocephalians Therocephalians of the Kotel'nich Tetrapod Assemblage Genus Species Notes Images GorynychusG. masyutinaeA wolf-sized therocephalian, the largest predator in the assemblageKarenitesK. ornamentatusA small, possibly insectivorous therocephalianKoksharoviaK. grechoviA small therocephalianKotelcephalonK. viatkensisMuchiaM. microdentaKnown only from a fragment of the lower jawPerplexisaurusP. foveatusA small, potentially semiaquatic therocephalianScalopodontesS. kotelnichiViatkosuchusV. suminiA large therocephalian
Gorgonopsians Gorgonopsians of the Kotel'nich Tetrapod Assemblage Genus Species Notes Images NochnitsaN. geminidensA very small, basal gorgonopsianViatkogorgonV. ivakhnenkoiA small basal gorgonopsian, suggested to be nocturnal
Anomodontians Anomodontians of the Kotel'nich Tetrapod Assemblage Genus Species Notes Images Dicynodontidaeindet.An indeterminate dicynodontid known from a partial skull.SuminiaS. getmanoviA small, tree-dwelling herbivore
Reptiles Reptiles of the Kotel'nich Tetrapod Assemblage Genus Species Notes Images DeltavjatiaD. rossicaA pareiasaur; the largest and most common tetrapod in the assemblageEmeroleterE. levisA small lizard-like nycteroleterid
References