List of Graphis (lichen) species

This is a list of species in the genus Graphis, a lichen-forming fungal genus in the family Graphidaceae. Species of Graphis are crustose lichens that form thin, often pale grey or greenish patches on bark (and less often rock), with elongated, slit-like fruiting bodies that trace dark, sometimes branching lines across the surface—an appearance that inspired the common name "script lichens". Graphis is widespread but especially diverse in tropical and subtropical regions, and it was established in 1763 by the French naturalist Michel Adanson.

, Species Fungorum (in the Catalogue of Life) accepts 300 species of Graphis. This article compiles those currently accepted species names; because the genus is actively revised, it is a snapshot that changes as names are synonymized, reinstated, or transferred between genera. Studies using molecular phylogenetics have shown that the traditional, broader use of Graphis grouped together more than one evolutionary lineage, leading to extensive reclassification within Graphidaceae. One major outcome has been the reinstatement of Allographa for a separate script lichen lineage (formalised in 2018), and other former Graphis species have also been reassigned to genera such as Carbacanthographis and Diorygma. Species totals therefore vary with the source consulted—for example, a global key published in 2009 accepted 330 species of Graphis and listed a further 205 names as synonyms—so any list reflects both the taxonomic scope and the date of the underlying classification.

A Graphis abapha Graphis acervulans Graphis alba Graphis albidofarinacea Graphis albidula Graphis albinula Graphis albissima Graphis albocarpa Graphis albocolpata Graphis alboglaucescens Graphis albothallina – India Graphis allugallenensis Graphis alpestris Graphis amaliana Graphis analoga Graphis andamanica Graphis anfractuosa Graphis anguilliformis Graphis annulata Graphis aperiens Graphis aperta Graphis apertella Graphis aphaneomicrospora Graphis aphanes Graphis apoda Graphis appendiculata Graphis arbusculiformis Graphis archeri Graphis arecae Graphis argentea Graphis assimilis Graphis astraea Graphis aterrima Graphis atroflava Graphis atrorubens Graphis atrosanguinea Graphis aurantiaca Graphis australosiamensis Graphis awaensis Graphis awasthii

B Graphis bakeri Graphis balaghatensis Graphis balansana Graphis batanensis Graphis beaumontii Graphis bernadetiae Graphis bicolor Graphis bicrenatula Graphis bipartita Graphis bogoriensis Graphis boliviana Graphis borealis Graphis botryosa Graphis bougainvilleae Graphis brachycarpa Graphis brahmanensis Graphis breussii Graphis brevicarpa Graphis brittoniae Graphis bulacana Graphis bungartzii

C

Graphis caelata Graphis caesiella Graphis caesiocarpa Graphis caesioglauca Graphis cambodiensis Graphis capillacea Graphis caribica Graphis carmenelisana Graphis castanopsidis Graphis catherinae Graphis celtidis Graphis centrifuga Graphis cervina Graphis cervinonigra Graphis chlorocarpella Graphis chlorotica Graphis chondroplaca Graphis chromothecia Graphis chungii Graphis cinnamomea Graphis cincta Graphis circumradians Graphis clavata Graphis coarctata Graphis coenensis Graphis colliculoides Graphis collinsiae Graphis compressa Graphis compulsa Graphis conferta Graphis confinis Graphis connectens Graphis consociata Graphis conturbata Graphis cooperta Graphis coriacea Graphis corrugata Graphis crassilabra Graphis crebra Graphis cremicolor Graphis crocea Graphis crystallifera Graphis cupei Graphis curvula Graphis cycadicola

D Graphis darjeelingensis Graphis decussata Graphis deformis Graphis deightonii Graphis delicatula Graphis dendrogramma Graphis dendroidea Graphis denudans Graphis descissa Graphis desquamescens Graphis detrita Graphis devestiens Graphis diaphoroides Graphis dicarpoides Graphis dichotoma Graphis discarpa Graphis dispersa Graphis distans Graphis distincta Graphis dracaenae Graphis dupaxana Graphis duplicata Graphis duplicatoinspersa

E

Graphis eburnea Graphis eimeoensis Graphis elegans Graphis elegantula Graphis elevata Graphis elongatoradians Graphis emersa Graphis emersella Graphis endophaea Graphis endoxantha Graphis enteroleuca Graphis epimelaena Graphis epiphloea Graphis erythraea Graphis erythrocardia Graphis eugeniae Graphis evirescens Graphis exalbata

F

Graphis fasciata Graphis ferruginea Graphis ferrugineodisca Graphis ficicola Graphis filiformis Graphis fissurata Graphis fissurinoides Graphis flabellans Graphis flavicans Graphis flavopalmicola Graphis flavovirens Graphis flexibilis Graphis flexuosa Graphis flindersiae Graphis flosculifera Graphis foliicola Graphis formosana Graphis fulminatrix Graphis fumosa Graphis furcata

G Graphis galactoderma Graphis geraensis Graphis gibberosa Graphis glaucella Graphis glaucescens Graphis glaucocaesia Graphis glaucocinerea Graphis glaucovirens Graphis gloriosensis Graphis gomphospora Graphis gonimica Graphis gracilenta Graphis gracilis Graphis graeffeana Graphis grammitica Graphis granulocarpa Graphis grossula Graphis guangdongensis Graphis guimarana Graphis guineensis

H Graphis haemolobata Graphis haleana Graphis halonata Graphis handelii Graphis hatschbachii Graphis heterocarpa Graphis hinnulea Graphis hochreutineri Graphis hodgesiana Graphis homographiza Graphis hongkongensis Graphis hunana Graphis hunanensis Graphis hyalinella Graphis hyphosa Graphis hypocrellina

I Graphis ignea Graphis illota Graphis immersa Graphis immersella Graphis immersicans Graphis implexula Graphis implicata Graphis inaequalis Graphis inamoena Graphis increbrior Graphis indica Graphis infida Graphis inopinata Graphis insculpta Graphis inspersa Graphis inspersolongula Graphis inspersonorstictica Graphis insulana Graphis insularis Graphis intermediella Graphis interversa Graphis intexta Graphis inustuloides Graphis inversa Graphis isidiza

J Graphis japonica Graphis jatrophae Graphis jeanmuelleri Graphis jejuensis Graphis jinhuana

K Graphis kakaduensis Graphis karstenii Graphis kavintuca Graphis kelungana Graphis kermesina Graphis khaojoneana Graphis khaoyaiensis Graphis kjellbergii Graphis kollaimalaiensis Graphis koltermaniae Graphis koratensis Graphis koreana Graphis kousyuensis Graphis kurokawae

L

Graphis laeviuscula Graphis latibasa Graphis lecanorina Graphis leioplaca Graphis leptaleocarpa Graphis leptocarpa Graphis leptocarpoides Graphis leptoclada Graphis leptotremoides Graphis librata Graphis limae Graphis lindsayana Graphis lineola Graphis litoralis Graphis longiramea Graphis lopingensis Graphis lucifica Graphis lueckingiana Graphis lueckingii Graphis luluensis Graphis lurizana Graphis lussuensis

M Graphis macrocarpa Graphis macularis Graphis maharashtrana Graphis makhijae – India Graphis malacoderma Graphis manhaviensis Graphis manipurensis Graphis maomingensis Graphis marginifera Graphis maritima Graphis marusae Graphis medusula Graphis meghalayensis Graphis mellis-insulae Graphis microsperma Graphis mikuraensis Graphis mindanaoensis Graphis mocimboensis Graphis modesta Graphis mokanarum Graphis moriensis – India Graphis moultonii Graphis multibrachiata Graphis murali-elegans Graphis muscicola Graphis myriocarpa Graphis myriocarpiza Graphis myriocarpoides Graphis myrtacea

N Graphis neeladriensis Graphis neglecta Graphis negrosina Graphis nematodes Graphis nematodiza Graphis neoelongata Graphis neoraensis Graphis nerurensis Graphis nigeriensis Graphis nigririmis Graphis nigrocarpa Graphis nigroglobosa Graphis nilgiriensis Graphis norfolkensis Graphis norsorediata Graphis norstictica Graphis noumeana Graphis nudanorsticta

O Graphis obtusior Graphis ocellata Graphis oligospora Graphis omiana Graphis ondensis Graphis overeemii Graphis oxyclada

P

Graphis pachysporella Graphis pallescens Graphis palmensis Graphis palmicola Graphis palmyrensis Graphis pananensis Graphis pandanicola Graphis panhalensis Graphis papillifera Graphis paradisserpens Graphis paradoxa Graphis paradussii Graphis paralleloides Graphis paranaensis Graphis paraschiffneri Graphis paraserpens Graphis pedata Graphis pernambucoradians Graphis persicina Graphis persoonii Graphis persulcata Graphis pertricosa Graphis pinicola Graphis planetocarpa Graphis platycarpella Graphis plumbeidisca Graphis plumierae Graphis polystriatosubmuriformis Graphis propinqua Graphis prunicola Graphis pseudoglyphis Graphis psidii Graphis puiggarii Graphis pulicaris Graphis pustulata Graphis pustulosa

R Graphis radians Graphis radiatoflexuosa Graphis rajapakshana Graphis ramosa Graphis redingeriana Graphis rigidula Graphis rimosothallina Graphis riopiedrensis Graphis robertusii Graphis rockii Graphis rondoniana Graphis rongklaensis Graphis rosae-emiliae Graphis rosalbinana Graphis roseotincta Graphis rubricosa

S

Graphis saipanensis Graphis salacinilongiramea Graphis sandalon Graphis santanderiana Graphis sapii Graphis saxicola Graphis saxiseda Graphis schiffneri Graphis schizogramma Graphis schizograpta Graphis schroederi Graphis schummiana Graphis scripta Graphis seawardii Graphis semiaperta Graphis separanda Graphis setschwanensis Graphis singaporina Graphis sirohiensis Graphis sitapurensis Graphis slendrae Graphis solmariana Graphis sororcula Graphis sphalera Graphis spodoplaca Graphis srilankensis Graphis stellata Graphis stenograpta Graphis stenospora Graphis stenotera Graphis stipitata Graphis streimannii Graphis striata Graphis subalbostriata Graphis subaltamirensis Graphis subangustata Graphis subcabbalistica Graphis subcontorta Graphis subcupei Graphis subcurva Graphis subelegans Graphis subelongata Graphis subfiliformis Graphis subhiascens Graphis subinsulana Graphis subintegra Graphis subintermedians Graphis subintricans Graphis sublitoralis Graphis submarginata Graphis subolivacea Graphis subparilis Graphis subpulicaris Graphis subregularis Graphis subrufula Graphis subserpentina Graphis subtecta Graphis subtenella Graphis subtracta Graphis subvittata Graphis sulphurella Graphis sundarbanensis Graphis supernata Graphis supracola Graphis suzanae Graphis syzygii

T Graphis taneina Graphis tapetica Graphis tenella Graphis tenellula Graphis tenoriensis Graphis tenuescens Graphis tenuirima Graphis tenuis Graphis tenuissima Graphis tetracarbonisata Graphis tetralocularis Graphis theae Graphis thoroldii Graphis thunsinhalayensis Graphis timidula Graphis tonglonensis Graphis tonkinensis Graphis treubii Graphis tricolor Graphis triquetra Graphis triseptata Graphis triticeella Graphis tsunodae Graphis tumida Graphis tumidulella Graphis turgidula

U Graphis umbrina Graphis urandrae

V Graphis valparaiensis Graphis venosa Graphis venusta Graphis vermifera Graphis verrucariiformis Graphis verrucata Graphis verruciformis Graphis verrucoserpens Graphis vinosa Graphis virens Graphis virescens Graphis viridithallina Graphis vittata Graphis vulvulescens

X Graphis xylophaga

Y Graphis yaucoensis Graphis yunnanensis

Z Graphis zollingeri Graphis zonatula

See also List of Allographa species

References