The Koshelevka Formation is a Permian formation in Russia containing the Tshekarda lagerstätte, which preserves over 200 insect species.
Insects
Holometabola
HolometabolansGenusSpeciesHigher taxonNotesImagesSylvacoleusS. richteri, S. sharoviTshekardocoleidaeHas a relatively long abdomen, named after the Sylva RiverSylvacoleodesS. admirandusTshekardocoleidaeResembles Sylvacoleus, but has a narrower headTshekardocoleusT. magnusTshekardocoleidaeType genus of the family, wing venation quite similar to the hypothetical ancestral beetleCavalarvaC. caudataHolometabola incertae sedisShares characteristics with larvae from numerous holometabolan orders
Paraneoptera ParaneopteransGenusSpeciesHigher taxonNotesImagesGraticladusG. apiatusIschnoneuridae (Hypoperlida)Placed in Hypoperlida, which was reclassified as stem-Paraneoptera in 2017TshekardoclardusT. sparsusTococladidaePlaced in Hypoperlida, which was reclassified as stem-Paraneoptera in 2017
Polyneoptera (excluding Notoptera) Non-Notopteran PolyneopteransGenusSpeciesHigher taxonNotesImagesArachnophasmaA. scurraPermophasmatidaeOldest stick insect known, has very long legs and a short bodyUraloblattaU. insignisArchimylacridae ("Eoblattodea")Has a relatively long oviscapt
Notoptera NotopteransGenusSpeciesHigher taxonNotesImagesArtinskaA. infigurabilisLemmatophoridaeBardapteronB. ovaleAtactophlebiidaeFormerly placed with paleodictyopterans, known from the less diverse Barda locality as opposed to ChekardaChelopterumC. ultimumChelopteridaeCucullistrigaC. cucullataIdelinellidaeNamed after the hood-like pronotum covering its headCuliciformaC. formosaLemmatophoridaeSimilar to MegorkhosaCzekardiaC. blattoidesEoblattida incertae sedisPreserves stripes and spots on its tergites, only known from a nymphDepressopterumD. bardumLiomopteridaeEuryptilodesE. commatulusEuryptilonidaeKnown from wings and exuviae of nymphsIblattaI. attrepidaEoblattida incertae sedisOnly known from a nymph, has quite short legs and a small headJubalaJ. pectinataEuryptilonidaeFormerly placed within SylvardembiaKhosarophlebiaK. sylvaensisLiomopteridaeFormerly known from a wing fragment, but had a complete specimen discovered laterKishertiaK. tricubitalisPinideliidaeDistinguished from Pinidelia by wing venation and sizeKoshelevkaK. megakhosaroidesDoubraviidaeFormerly included in CerasopterumKungurocaudaK. spinosaSylviaphlebiidaeSecond-smallest member of the family behind SylvophenopteraKunguroliomaK. cancellataLiomopteridaeLemmatonymphaL. gracilis, L. gracilissimaLemmatophoridaeIdentifiable to family based on its tarsi and cerciLiomopterellaL. kunguricaLiomopteridaeMicaideliaM. minutissimaIdeliidaeSimilar to AnaideliaNeprotembiaN. truncataPermulidaeSimilar to NeraphidiaNeraphidiaN. mitis, N. rigidaNeraphidiidaeFormerly only known from part of a forewing, but a complete specimen was discovered laterNovokshonovusN. ignoratusAtactophlebiidaeRelated to KirkorellaOrnaticostaO. novokshonoviLiomopteridaeParakhosaraP. coalitaMegakhosaridaeParasheimiaP. truncata, P. rotundataSheimiidaeLarger than Sheimia and has a wider pronotumParastenaropoditesP. stirpsMesorthopteridaeParasylvaellaP. umbra, P. minorLiomopteridaeParasylviodesP. tetracladusLiomopteridaeParatillyardembiaP. sepicolorataProtembiidaeHas an unusual pronotumParmapteraP. permianaParmapteridaePlaced within its own monotypic familyPectinokhosaraP. sylvardembiodesMegakhosaridaeHas elongated wingsPermedaxP. effertusTillyardembiidaeDiffers from all other known grylloblattid nymphs by its elongate body, but this is also a character of adult tillyardembiidsPermulaP. tshekardensis, P. minorPermulidaeFormerly placed within Sojanopermula, with Permula itself being a replacement name for Allicula.PermoshurabiaP. kunguricaGeinitziidaeOne of the earliest geinitziidsPermostrigaP. augustalisIdelinellidaePseudosheimiaP. caudataSheimiidaeHas longer cerci than SheimiaScutistrigaS. scutataIdelinellidaeNamed after its shield-like pronotumSheimiaS. tshekardensisSheimiidaeBears larger eyes than other Sheimia speciesStrigullaS. cuculiophorisIdelinellidaeFormerly placed within EuryptilonSuksunusS. bicodexCacurgidaeSimilar to KochopteronSylvaclinicusS. echinatusGrylloblattida incertae sedisOnly known from nymphsSylvafossorS. forcipatusProbnidaeHas unusual earwig-like cerciSylvamicropteronS. harpaxGrylloblattida incertae sedisAn unusual micropterous grylloblattidSylvaphlebiaS. tuberculata, S. fucataSylvaphlebiidaeHas a pigmented wing stripSylvapriscaS. gravis, S. forta, S. focaleataLemmatophoridaeNamed after the lemmatophorid genus ParapriscaSylvardembiaS. matura, S. tamaenaSylvardembiidaeSylvastrigaS. mirandaIdelinellidaeSylvictorS. majorLiomopteridaeOne of the largest notopterans from KoshelevkaSylviodesS. perloidesMesorthopteridaeFormerly placed in IdeliidaeSylvonymphaS. tshekardensisGrylloblattida incertae sedisUnusually for a grylloblattid, the larva has gillsSylvophenopteraS. perlongata, S. fimbriataSylvaphlebiidaeResembles Phenopterum but has a longer ovipositorTillyardembiaT. antennaeplana, T. ravisedorumTillyardembiidaeMost common insects from Koshelevka alongside the earliest insects with preserved pollen attached to themTshekardanymphaT. lienterica, T. bardensisGrylloblattida incertae sedisSplit from TshekarditesTshekardeigmaT. rasnitsyniEpideigmatidaeTshekardeliaT. mediaIdeliidaeOne of the first ideliids known from a well-preserved bodyTshekarditesT. comitialisGrylloblattida incertae sedisFirst established as a nymphal form taxon, before all other species were split outTshekardominaT. maculatus, T. imbecilla, T. imbecillissima, T. subincurvataTshekardominidae (Grylloblattida)Type specimen preserves several eggs inside the abdomen, rest of specimens are all female (possibly suggesting flightless males?)TshekardophlebiaT. capitataSylvaphlebiidaeResembles Sylvaphlebia but has a larger headTshekardushkaT. artenatisReculida incertae sedisOnly known from a nymph. Preserves an oval impression, likely representing a bolus of inorganic materialUraliomaU. maximaLiomopteridaeIncompletely preserved, may belong to a separate genusUralopriscaU. uralicaLemmatophoridaeFormerly included in ParapriscaVisheriferaV. sylvaensisVisheriferidaeOriginally described from a late Permian locality in the Solikamsk Formation
Misc. Arthropoda
Miscellaneous arthropodsGenusSpeciesHigher taxonNotesImagesPermarachneP. novokshonoviUraraneidaPreviously identified as a spiderPermocrassacusP. novokshonoviScolopendromorphaFirst described Permian centipede
Plants
Seed plants
Conifers
ConifersGenusSpeciesHigher taxonNotesImagesCyparissidiumC. appressumVoltzialesFormerly placed within WalchiaKungurodendronK. sharoviiVoltzialesSimilar to Walchia, although differs in having seed stalksTaxodiellaT. bardaeanaVoltzialesFormerly placed within Walchia, synonymous with several species
References