The Kladno Formation is a Carboniferous formation in the Czech Republic which preserves a wide array of temnospondyls, lepospondyls and fish, among other fauna.
Paleobiota
Invertebrates
InvertebratesGenusSpeciesHigher taxonNotesImagesNecymylacrisN. scudderiBlattodeaFormerly included within PhylloblattaBohemiatupusB. elegansMeganeuridaeOne of the larger meganisopteransBojophlebiaB. prokopiPalaeopteraLargest insect by wingspan other than palaeodictyopterans or griffinflies.CarbotripluraC. kukalovaeDicondyliaSister group to Pterygota, formerly classed as a Bojophlebia nymph.PronaiditesP. carbonarius, P. arenivorusTubificidae"P. crenulatus" actually an ichnofossilMicroconchusM. vorax, M. minimusTentaculitaFormerly classed as a spirorbidProlimulusP. woodwardiBellinuridaeLikely clades with other genera like Alanops.EolycosaE. lorenziiMesothelae incertae sedisSpider affinities doubted in a 1996 paperGeralycosaG. friciiArthromygalidaeMay not be a spiderRakovniciaR. antiquaTetrapulmonata incertae sedisPyritaraneaP. tubiferaAraneae incertae sedisSynonymised with EopholcusNyranytarbusN. hofmanni, N. longipesTrigonotarbidaFormerly placed within HemiphrynusDoubravatarbusD. kraftiAphantomartidaeLikely arboreal, due to being found on a liana-like plant and having relatively gracile legs.TynecotarbusT. tichavekiTrigonotarbidaCollected alongside two more fragmentary indeterminate fossilsPalaraneaP. borassifoliaAraneae incertae sedisKnown from a different locality to other Kladno faunaAnthracomartusA. carcinoides, A. bohemica, A. elegansTrigonotarbidaIsobuthusI. nyranensisScorpionesProthelyphonusP. bohemicusUropygiSynonymous with GeralinuraDinopilioD. gigasArachnida incertae sedisVery large fossil, hence the nameGeratarbusG. bohemicusPhalangiotarbidaOrthotarbusO. nyranensisPhalangiotarbidaPleurolycosaP. proliferaArthropoda incertae sedisNyraniusN. costulatus, N. tabulatusXyloiuloideaFormerly within XylobiusPleurojulusP. biornatus, P. levisPleurojulidaAlso known from Mazon CreekIsojulusI. constansPleurojulidaSynonymised with two Pleurojulus speciesPurkyniaP. lataZosterogrammidaVery poorly preserved
Vertebrates
VertebratesGenusSpeciesHigher taxonNotesImagesArchaeothyrisA. spOphiacodontidaeOnly known from a jaw fragmentBrouffiaB. orientalisEureptiliaFormerly placed within GephyrostegusCoelostegusC. prothalesEureptiliaRecovered as the basalmost eureptileGephyrostegusG. bohemicusJaekel, O. (1902): Gephyrostegus bohemicus, n.g. n.sp. Zeitschrift der Deutschen geologischen Gesellschaft 54: (Verhandlungen der Gesellschaft): 127–132GephyrostegidaeFormerly included another species, G. watsoniSolenodonsaurusS. janenschiReptiliomorpha?Resembles diadectomorphs, yet likely not closely related to amniotesUrocordylusU. angularisUrocordylinaeAlso known from IrelandSauropleuraS. scalarisSauropleurinaeFormerly included in UrocordylusScincosaurusS. crassusScincosauridaeFormerly synonymised with Keraterpeton, related to diplocaulidsKeraterpetonK. galvaniDiplocaulidaeBore an unusually long tailHyloplesionH. longicostatum"Microsauria"Synonymous with "Orthocosta" and "Seeleya"MicrobrachisM. pelikaniRecumbirostraSynonymised with three other species in the same genusRicnodonR. copeiRecumbirostra?May be related to SaxonerpetonCrinodonC. limnophyesTuditanidaeName derives from an anagram of RicnodonSparodusS. validusGymnarthridae?Resembles HylerpetonDiplovertebronD. punctatumEmbolomeriSynonymous with "Nummulosaurus"CapetusC. palustrisTemnospondyliOne of the most basal temnospondylsCochleosaurusC. bohemicusCochleosauridaeAdult specimens only found in 2003MordexM. calliprepesTrematopidaeUnclear position within the familyMattauschiaM. laticepsTrematopidaeFormerly included in Mordex, includes "Potamochoston" and several "Limnerpeton" speciesNyranerpetonN. amilneriMicromelerpetontidaeAlso known from the Montceau-les-Mines lagerstätteLimnogyrinusL. elegansMicromelerpetontidaeFormerly included within "Limnerpeton"OestocephalusO. granulosum, O. nanumAistopodaFormerly included within OphiderpetonPhlegethontiaP. longissima, "Dolichosoma scutiferum"AistopodaFormerly included within the nomen nudum "Dolichosoma"BaphetesB. orientalisBaphetidaeFormerly known as "Loxomma bohemicum"StambergichthysS. macrodensActinopterygiiLikely a high-level predator in its ecosystemSceletophorusS. biserialis, S. verrucosusTrissolepididaePyritocephalusP. sculptusHaplolepiformesSynonymous with other Pyritocephalus species
Plants
Non-seed plants
Non-seed plantsGenusSpeciesHigher taxonNotesImagesKladnostrobusK. clealii, K. psendaeLycopsidaWithin the monotypic family Kladnostrobaceae due to differing sporangia structureLepidophloiosL. acerosusLepidodendralesMistakenly spelled "Lepidofloios" in the paper reporting its presenceLepidodendronL. acutum, L. simile, L. aculeatum, L. ophiurum, L. lycopodioidesLepidodendralesOne of the most abundant Carboniferous plantsFlemingitesF. spLepidodendralesCone taxonLepidocarponL. majusLepidodendralesSporangium taxonSigillariaS. rugosa, S. diplodermaLepidodendralesS. diploderma only known from a small fragmentSyringodendronS. spLepidodendralesStem taxonOmphalophloiosO. feistmantelliIsoetalesQuestionable specimen known from a fragment of a fertile axisStigmariaS. ficoidesLepidodendralesForm taxon representing rootsCalamitesC. carinatus, C. distachya?, C. cistii?CalamitaceaeSecond species is closest to C. distachya, but poor preservation means its affinity is unclearAnnulariaA. radiataCalamitaceaeLeaf taxon of Calamites stemsAsterophylitesA. longifolius, A. equisetiformis, A. grandisCalamitaceaeOne specimen preserves the tip of a branch with dense leavesSphenophyllumS. cuneifolium, S. pseudoaquense, S. ?majus, S. priveticenseSphenophyllalesVery common, but often poorly preservedBowmanitesB. priveticensis, B. myriophyllus, B. pseudoaquensis, B. brasensisSphenophyllalesCones of SphenophyllumLobatopterisL. aspidioidesMarattialesOnly one small fragment preservedCorynepterisC. sternbergii, C. angustissima, C. essinghiiZygopteridalesC. sternbergii may be synonymous with C. angustissimaSenftenbergiaS. plumosaTedeleaceae (Filicales)Common elsewhere, yet rare in KladnoZeilleriaZ. delicatulaFilicales incertae sedisShares its name with a Jurassic brachiopodRenaultiaR. crepiniiFilicales incertae sedisOnly known from a fragmentSphenopterisS. rotundifolia, S. mixta, S. delicatula, S. acutiloba, S. ujezdensis, S. pulcherrima, S. cirrhifolia, S. spinosaFilicales incertae sedis (rotundifolia), Lyginopteridales (others)Includes seed plants and a fernAphlebiaA. spIndeterminateArtificial taxon; may belong to another species preserved.NoeggerathiaN. foliosa, N. intermediaNoeggerathialesMost common plant in the sampled fossilsArchaeonoeggerathiaA. schatzlarensisNoeggerathialesDiscosoropterisD. zlatkokvacekii, D. chlupatumFilicales incertae sedisLikely a short, upright plantSpenceritesS. leismanii, S. havlenaeLycopsida incertae sedisA member of the "sub-arborescent" lycophytesDendraenaD. pinnatilobataAnachoropteridaceae (Filicales)Likely grew near river margins, a very common plantKidstoniaK. heracleensisFilicalesLikely a small understorey fernDesmopterisD. alethopteroides, D. longifoliaFilicalesLikely bore procumbent (growing along the ground) rhizomesSelaginellaS. gutbieri, S. labutaeSelaginellaceaeExtant genusPalaeostachyaP. distachya, P. gracilimaCalamitaceaeCone taxon, although with a complete plant known from the early Permian of ChinaPecopterisP. aspidioides, P. tuffiticaPsaroniaceaeLikely a mid-sized tree fernSonapterisS. pilsensis, S. barthelii, S. bekiiBotryopteridaceae (Filicales)Possibly a liana-like plant?OligocarpiaO. lindsaeoidesGleicheniales?Probably a liana-like plantAdiantitesA. spProgymnospermopsidaLikely a low-lying plantEchinosporangitesE. libertiteFilicopsida?Sporangium taxon, parent plant unknownPolysporiaP. drabekii, P. rothwelii, P. radvanicensis, P. robustaIsoetalesA sub-arborescent lycophyteHuttoniaH. spicataEquisetalesCone taxonThomasitesT. serrata, T. elongatusLycopsidaT. elongatus formerly placed in Selaginellites
Seed plants
Seed plantsGenusSpeciesHigher taxonNotesImagesAlethopterisA. lonchitica, A. distantinervosa, A. pseudograndinioides, A. kettneri, A. serliiMedullosalesA. pseudograndinioides is a replacement name for the illegitimate "A. grandinioides"NeuromariopterisN. scandensCallistophytalesLikely a creeping plant growing on ridge slopesEusphenopterisE. nummulariaLyginopteridalesWhile a large (>1 m) frond was found, it could not be transported from the site.LonchopterisL. rugosaMedullosalesPossible record of L. chandesrisii (although based on a poorly preserved specimen) also knownMacroneuropterisM. scheuchzeriMedullosalesFormerly placed within NeuropterisNeuropterisN. plicataMedullosalesPlacement of this species is unclear in relation to N. ovata.OdontopterisO. reichianaMedullosalesMay be synonymous with O. brardiiCallipteridiumC. rubescens, C. armasiiMedullosalesMay be synonymous with C. jongmansiiLaveineopterisL. loshii, L. tenuifolia, L. bohemica, L. lubnensis, L. hollandicaMedullosalesL. loshii is abundant elsewhere, while some other species are endemic to the Kladno FormationPalaeoweichseliaP. defranceiMedullosalesDescribed from Kladno in a 2020 overview paper despite specimens being already known priorLinopterisL. neuropteroides, L. obliqua, L. weigeliiMedullosalesL. neuropteroides is split into two forms, neuropteroides and minor.ParipterisP. linguaefoliaMedullosalesIncluded alleged records of P. giganteaHavlenaeaH. coriacea, H. stradonitzensisMedullosalesFormerly placed within Neuropteris Mixoneura M. muensterifolia Medullosales Too poorly preserved to place within a new genus, yet does not belong to Mixoneura either RhacopterisR. elegansPteridospermatophytaOnly known from one isolated pinnule (leaf cluster)CordaitesC. theodorii, C. wartmannii, C. kladnoensis, C. borassifoliusCordaitalesC. theodorii is only known from this formationPalmatopterisP. furcataLyginopteridalesProbably a liana-like plantMariopterisM. muricataLyginopteridalesLikely a vining plant, as its fossils are all associated with Lepidodendron.FortopterisF. radnicensisMedullosalesNot given a type specimen in its original description within Mariopteris.RhodeitesR. gutbieriLyginopteridalesLikely a canopy-dwelling climber
References