This article records new taxa of fossil plants that were described during the year 2019, as well as other significant discoveries and events related to paleobotany that occurred in 2019.

Mosses Name Novelty Status Authors Age Type locality Location Notes ImagesHeinrichsiellaGen. et sp. novValidBippus et al.JurassicA moss, possibly related to the family Polytrichaceae or Timmiellaceae. Genus includes new species H. patagonica.KulindobryumGen. et sp. novValidIgnatov in Mamontov & IgnatovMiddle or Late JurassicUkureyskaya Formation()A form genus of dispersed moss capsules. Genus includes new species K. taylorioides.Polycingulatisporites multiverrucataSp. novIn pressSantamarina in Santamarina et al.Late Cretaceous (Cenomanian)Mata Amarilla FormationSpores of a member of Bryophyta of uncertain phylogenetic placement, possibly of sphagnaceous affinity. Announced in 2019; the final version was scheduled to be published in 2020.Sphagnum heinrichsiiSp. novValidIgnatov et al.Late EoceneRovno amberA moss, a species of Sphagnum.PaleaethallusGen. et sp. novValidMamontov, Katagiri & Borovich in Mamontov & IgnatovLate JurassicGlushkovo Formation()A thalloid bryophyte. Genus includes new species P. squarrosus.

Liverworts Name Novelty Status Authors Age Type locality Location Notes ImagesKhasurtythallusGen. et sp. novValidMamontov in Mamontov & IgnatovEarly Cretaceous()A Marchantiidae liverwort. The type species is K. monosolenioides.Ricciopsis sandaolingensisSp. novValidLi & Sun in Li et al.Middle JurassicXishanyao FormationA Ricciaceae liverwort.Thallites yangcaogouensisSp. novValidWang et al.Late TriassicYangcaogou FormationA plant of uncertain phylogenetic placement, probably a liverwort.

Ferns and fern allies Name Novelty Status Authors Age Type locality Location Notes ImagesAlloiopteris loecseiSp. novValidPšenička et al.Carboniferous (Moscovian)A zygopterid fern.Annularia noronhaiSp. novValidCorreia et al.Carboniferous (Gzhelian)Douro BasinA member of the family Calamitaceae. Announced in 2019; the final version of the article naming it was published in 2021. Azolla keujaSp. novValidJud, De Benedetti, Gandolfo & HermsenPaleocene (Danian)Salamanca FormationA species of Azolla.BerendtiopterisGen. et comb. novValidSadowski et al.EoceneBaltic amberEurope (Baltic Sea region)A plant of uncertain phylogenetic placement, probably a fern; a new genus for "Pecopteris" humboldtiana. BifariusothecaGen. et sp. novValidX.H.Zhao ex DoweldLate PermianLongtan FormationA marattialean fern. Genus includes new species B. notocathaysica Doweld.Bowmanites yongchangensisSp. novValidSun et al.Permian (Cisuralian)A member of Sphenophyllales.Clavatisporites cenomanianaSp. novIn pressSantamarina in Santamarina et al.Late Cretaceous (Cenomanian)Mata Amarilla FormationSpores of a member of Filicopsida of uncertain phylogenetic placement. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020.Collarisporites minorSp. novIn pressSantamarina in Santamarina et al.Late Cretaceous (Cenomanian)Mata Amarilla FormationSpores of a member of Filicopsida of uncertain phylogenetic placement. Announced in 2019; the final version of the article naming it is scheduled to be published in 2020.Cyathocarpus yongchangensisSp. novValidSun & Sun in Sun et al.Permian (Cisuralian)Shanxi FormationA fern related to Psaronius.FlorathecaGen. et sp. novValidLundgren et al.Early PermianRío Genoa FormationA member of Marattiales. Genus includes new species F. apokalyptika.Germera brousmicheaeSp. novValidÁlvarez-VázquezCarboniferous (Westphalian)A member of Filicopsida.Hausmannia olaensisSp. novValidGolovneva & GrabovskiyLate Cretaceous (Santonian–early Campanian)A Dipteridaceae umbrella fern.HeinrichsiaGen. et sp. novValidRegalado et al.CretaceousBurmese amberA fern belonging to the family Pteridaceae. Genus includes new species H. cheilanthoides.KamathecaGen. et comb. novValidDoweldPermianA marattialean fern; a new genus for "Acitheca" gigantea Esaulova.Marsilea sprungerorumSp. novValidHermsenEoceneGreen River FormationParachute Creek MemberA Marsilea species water fern.NeolobatannulariaGen. et sp. novValidSun & Li in Wang et al.Late TriassicYangcaogou FormationA member of Equisetales. Genus includes new species N. liaoningensis.Osmundastrum gvozdevaeSp. novValidBazhenova & BazhenovMiddle Jurassic (Bathonian)()A species of Osmundastrum.Palaeosorum waipiataSp. novValidKaulfuss et al.Early MioceneA member of the family Polypodiaceae.Phlebopteris kirchneriSp. novValidBarbacka & Kustatscher in Barbacka, Kustatscher & BodorEarly Jurassic (Hettangian)Mecsek Coal FormationA fern belonging to the family Matoniaceae.Plenasium xieiSp. novValidCheng et al.Cretaceous A member of Osmundaceae. Announced in 2019; the final version of the article naming it was published in 2021.Polymorphopteris magdalenaeSp. novValidR.H.Wagner ex DoweldLate CarboniferousA marattialean fern.Polymorphopteris wagneriSp. novValidDoweldLate Carboniferous (Kasimovian)A marattialean fern.Polypodiisporites serratusSp. novValidD'Apolito et al.Pliocene–PleistoceneFossil spore of a member of the family Polypodiaceae.Polypodiisporites timidusSp. novValidD'Apolito et al.Pliocene–PleistoceneFossil spore of a member of the family Polypodiaceae.RinistachyaGen. et sp. novValidPrestianni & GessDevonian (Famennian)Witpoort FormationA member of Sphenophyllales. The type species is R. hilleri.RothwellopterisGen. et sp. novValidHe et al.Late PermianXuanwei FormationA fern belonging to the group Marattiales. Genus includes new species R. pecopteroides.Scolecopteris liberaSp. novValidLi et al.Permian (Asselian)Taiyuan FormationA marattialean fernScolecopteris renaultiiSp. novValidDoweldPermian (Cisuralian)A marattialean fern.Tiania resinusSp. novValidHe & WangPermian (Lopingian)Xuanwei FormationA member of Osmundales belonging to the extinct family Guaireaceae.

Lycophytes Name Novelty Status Authors Age Type locality Location Notes ImagesBergeria wenquanensisSp. novValidFeng, D'Rozario & ZhangCarboniferous (Viséan)Akeshake FormationA member of Lepidodendrales belonging to the family Flemingitaceae.GuangdedendronGen. et sp. novWang et al.Devonian (Famennian)Wutong FormationA member of Isoetales belonging to the group Dichostrobiles. Genus includes new species G. micrum.Omphalophloios wagneriSp. novValidOpluštil, Pšenička & BekCarboniferous (Moscovian)Illinois Basin()Sawdonia hippothecaSp. novValidBerry & GenselDevonian (probably late Givetian)Campo Chico FormationA member of Zosterophyllopsida.

Conifers Araucariaceae Name Novelty Status Authors Age Type locality Location Notes ImagesAgathoxylon giliiSp. novValidRíos-Santos & Cevallos-FerrizLate JurassicTodos Santos FormationAgathoxylon hoodiiComb novvalid(Tidwell & Medlyn) Gee et alLate JurassicMorrison FormationAn araucariaceous petrified wood.Moved from Araucarioxylon hoodii (1993)Agathoxylon jericonseSp. novValidRíos-Santos & Cevallos-FerrizLate JurassicTodos Santos FormationAgathoxylon kotaenseSp. novIn pressChinnappa, Rajanikanth & Pauline Sabina?Late Jurassic – Early CretaceousKota FormationA member of the family Araucariaceae.Agathoxylon parrensisSp. novValidRíos-Santos & Cevallos-FerrizPaleoceneLas Encinas FormationAraucaria balfourensisSp. novValidHill et al.CenozoicA species of Araucaria.Araucaria macrophyllaSp. novValidHill et al.CenozoicA species of Araucaria.Araucaria mollifoliaSp. novValidHill et al.CenozoicA species of Araucaria.Araucaria rothwelliiSp. novValidKvaček in Kvaček et al.Late Cretaceous (Campanian-Maastrichtian)Bozova FormationA species of Araucaria.Brachyphyllum garciarumSp. novValidCarrizo et al.Early Cretaceous (early Hauterivian/early Barremian)Springhill FormationProbably a member of the family Araucariaceae.

Cupressaceae Name Novelty Status Authors Age Type locality Location Notes ImagesAustrocupressinoxylonGen. et sp. novValidNunes et al.Early CretaceousA member of Cupressaceae. Genus includes new species A. barcinense.Austrohamia asfaltensisSp. novValidContreras et al.Early JurassicCañadón Asfalto FormationA member of the family Cupressaceae.Callitris blackburniiSp. novValidPaull et al.Middle MioceneA species of Callitris.Cupressinoxylon pliocenicaSp. novValidAkkemikPlioceneÖrencik FormationA member of the family Cupressaceae described on the basis of fossil wood.Mesocyparis sinicaSp. novValidCui et al.Paleocene (Danian)Wuyun FormationA member of the family Cupressaceae.Protaxodioxylon sahniiSp. novValidChinnappa, Kavali & RajanikanthLate Jurassic to Early CretaceousKota FormationA member of Cupressaceae, possibly related to Taxodium.Protodammara reimatamorioriSp. novValidMays & Cantrill Late Cretaceous (Cenomanian)Tupuangi FormationA member of Cupressaceae.Taxodioxylon cabullensisSp. novValidRíos-Santos & Cevallos-FerrizLate CretaceousPackard FormationA cupressaceous fossil wood.Taxodium viligenseSp. novValidGolovnevaLate Cretaceous (Coniacian)Chingandzha FormationA species of Taxodium.

Pinceae Name Novelty Status Authors Age Type locality Location Notes ImagesAbies cuitlahuaciiSp. novValidCevallos-Ferriz, Ríos-Santos & Lozano-GarcíaPleistoceneA fir.Pinus plioarmandiiSp. novValidAn et al.PlioceneA pine.Pinuxylon alonissianumSp. novValidMantzouka & Sakala in Mantzouka et al.Early MioceneA member of the family Pinaceae described on the basis of fossil wood.Schizolepidopsis borealisSp. novValidDomogatskaya & HermanEarly Cretaceous (Albian)Balyktakh FormationA member of the family Pinaceae.

Podocarpaceae Name Novelty Status Authors Age Type locality Location Notes ImagesDacrycarpus guipingensisSp. novValidWu et al.MioceneErzitang FormationA species of Dacrycarpus. Announced in 2019; the final version of the article naming it was published in 2021. KirketapelGen. et sp. novValidAndruchow-Colombo et al.Paleocene (early Danian)Salamanca FormationA member of the family Podocarpaceae. The type species is K. salamanquensis.Podocarpus pliomacrophyllusSp. novIn pressChen et al.Early Pliocene A species of Podocarpus. Announced in 2019; the final version of the article naming it is not published yet.

Other conifers Name Novelty Status Authors Age Type locality Location Notes ImagesCephalotaxus maguanensisSp. novValidZhang et al.Middle MioceneA species of Cephalotaxus.Cupressinocladus shelikhoviiSp. novValidGolovnevaLate Cretaceous (Coniacian)Chingandzha FormationA cheirolepidiaceous speciesFrenelopsis justaeSp. novValidBarral et al.Early Cretaceous (Albian)Escucha FormationA member of the family Cheirolepidiaceae.Ningxiaites shitanjingensisSp. novValidWei et al.Permian (Changhsingian)Sunjiagou FormationA conifer wood.Protocupressinoxylon carrizalenseSp. novValidCorrea et al.Late TriassicCarrizal Formation

Other seed plants Name Novelty Status Authors Age Type locality Location Notes ImagesAmyelon bogdenseSp. novValidWan, Yang & WangLate Permian or Early TriassicGuodikeng FormationA silicified gymnospermous root.ArazedispermumGen. et sp. novValidFriis, Crane & PedersenEarly Cretaceous (late Aptian-early Albian)Figueira da Foz FormationA seed plant belonging to the informal grouping Bennettitales-Erdtmanithecales-Gnetales. Genus includes new species A. lustanicum.AxsmithiaGen. et comb. novValidAnderson et al.TriassicAntarcticaA seed fern. Genus includes "Umkomasia" uniramia Axsmith et al. (2000).Bowenia johnsoniiSp. novValidHill et al.Early EoceneA cycad, a species of Bowenia.BrinkiaGen. et 2 sp. novValidKustatscher, Visscher & van Konijnenburg-van CittertPermian (Lopingian)Bellerophon Formation