Trabutina mannipara, or mana scale, is a species of mealybug found in the Middle East and southern Europe.CABI Invasive Species Compendium.Yair Ben-Dov (1988). "Manna scale, Trabutina mannipara (Hemprich & Ehrenberg) (Homoptera: Coccoidea: Pseudococcidae)". Systematic Entomology. Volume 13, Issue 4. It is the most well-known of the five species in the genus Trabutina, of which it is the type species,D.J. Williams and P. J. Gullan (2010). "Family-group names proposed in the family Pseudococcidae (Hemiptera: Sternorrhyncha: Coccoidea)".Zootaxa. due to its association with the biblical story of manna.Evelyna M. Danzig and Douglass R. Miller (1996). "A Systematic Revision of the Mealybug Genus Trabutina (Homoptera: Coccoidea: Pseudococcidae)." Israel Journal of Entomology, XXX. p. 7. T. mannipara feeds parasitically on tamarisk trees, and excretes a sweet substance which is sometimes collected for human consumption. Obsolete terms for it include Coccus manniparus and Trabutina palestina. This species can be found in the Sinai and Iraq.
It was described in 1829 by Hemprich and Ehrenberg (as Coccus manniparus),Yair Ben-Dov (1988). "Manna scale, Trabutina mannipara (Hemprich & Ehrenberg) (Homoptera: Coccoidea: Pseudococcidae)". Systematic Entomology. Volume 13, Issue 4. p. 388, 389. after investigation in the Sinai mountains.M. Bertholet (1862). "On the Manna of Sinai, and the Manna of Syria". The Pharmaceutical Journal and Transactions. Second Series, Volume 3. p. 274. Ehrenberg believed that the bite wounds in the tamarisk plant created as the insects fed on the plant caused a sweet sticky substance known locally as "manna", and which Ehrenberg associated with the biblical manna, to flow out of the plant. In 1929, F. S. Bodenheimer found that the manna was in fact produced by the insects themselves, and argued that the description of the sticky substance and its geographical region argued in favor of Ehrenberg's identification with the biblical manna. The identification with biblical manna continues to appear in more recent literature.
T. mannipara, like other mealybugs, is parasitic on plants, and in its excretions is a great deal of undigested material. In the case of mannipara, these excretions are high in sugar.
In the United States, where tamarisks are invasive species, T. mannipara has been tested for possible use in controlling tamarisks.
Notes
References