These species belong to Maladera, a genus of May beetles and junebugs in the family Scarabaeidae.
Maladera species
Amaladera Reitter, 1896 Maladera cobosi Baraud, 1964 Maladera espagnoli Baraud, 1964 Maladera euphorbiae (Burmeister, 1855) Maladera golovjankoi Medvedev, 1952 Maladera longipennis (Verdu, Mico & Galante, 1997) Aserica Lewis, 1895 Maladera secreta (Brenske, 1898) Maladera shaowuensis Ahrens, Fabrizi & Liu, 2021 Cycloserica Reitter, 1896 Maladera arenicola (Solsky, 1876) Maladera caspia (Faldermann, 1836) Maladera excisiceps (Reitter, 1896) Maladera kerleyi Ahrens, 2004 Maladera mechiana Ahrens, 2004 Maladera paraquinquidens Ahrens, 2004 Maladera quinquidens (Brenske, 1896) Maladera ustinovi (Gusakov, 2021) Eumaladera Nomura, 1967 Maladera fencli Ahrens, Fabrizi & Liu, 2021 Maladera fushanica (Ahrens, 2002) Maladera gravida Ahrens, 2006 Maladera latitibia Nomura, 1974 Maladera loi (Kobayashi, 1991) Maladera nitidiceps Nomura, 1967 Maladera nitididorsis Nomura, 1967 Maladera opaciventris (Moser, 1915) Maladera paralatitibia (Ahrens, 2002) Maladera planiuscula Nomura, 1974 Maladera puliensis (Ahrens, 2002) Maladera robustula Ahrens, Fabrizi & Liu, 2021 Maladera subrugata (Moser, 1926) Maladera wipfleri Ahrens, Fabrizi & Liu, 2021 Maladera yonaguniensis Kobayashi, Kusui & Imasaka, 2006 Eusericula Reitter, 1902 Maladera ahrensi Keith, 2005 Maladera farsensis Petrovitz, 1980 Maladera frischi Keith, 2011 Maladera girardi Keith & Ahrens, 2002 Maladera hodkovae Nikodým & Král, 1998 Maladera jiraskovae Sehnal, 2008 Maladera kermanensis Fabrizi, Rößner & Ahrens, 2018 Maladera mirzayansi Montreuil & Keith, 2009 Maladera modesta (Fairmaire, 1881) Maladera mofidii Montreuil & Keith, 2009 Maladera schnitteri Ahrens, Fabrizi & Rössner, 2016 Maladera sinaica Ahrens, 2000 Hemiserica Brenske, 1894 Maladera armipes (Arrow, 1945) Maladera ballariensis Ahrens & Fabrizi, 2016 Maladera bilobata (Arrow, 1945) Maladera brevis (Blanchard, 1850) Maladera eshanensis Ahrens, Fabrizi & Liu, 2021 Maladera fangchengensis Ahrens, Fabrizi & Liu, 2021 Maladera hsui Ahrens, Fabrizi & Liu, 2021 Maladera lorenzi Ahrens & Fabrizi, 2016 Maladera mutabilis (Fabricius, 1775) Maladera nasuta (Brenske, 1894) Maladera nasutella (Ahrens, 2004) Maladera rufopaca Ahrens, Fabrizi & Liu, 2021 Maladera severini (Brenske, 1896) Maladera subinaequalis Sreedevi & Ahrens, 2025 Maladera trilobata (Khan & Ghai, 1980) Macroserica Medvedev, 1952 Maladera allemandi Keith, 1998 Maladera angusta Baraud, 1990 Maladera apfelbecki Petrovitz, 1969 Maladera armeniaca Reitter, 1896 Maladera attaliensis Petrovitz, 1969 Maladera baluchestanica Petrovitz, 1971 Maladera besucheti Baraud, 1990 Maladera bruschii Sabatinelli, 1977 Maladera cerrutii Sabatinelli, 1977 Maladera dentipenis Sehnal & Simandl, 2008 Maladera fusconitens (Fairmaire, 1892) Maladera iraqensis Keith, 2000 Maladera kermanica Montreuil, 2016 Maladera khuzestanica Montreuil, 2016 Maladera krueperi Petrovitz, 1969 Maladera lodosi Baraud, 1975 Maladera loebli Baraud, 1990 Maladera lyciensis Petrovitz, 1969 Maladera miliouensis Keith & Miessen, 2009 Maladera ollivieri Keith, 1998 Maladera phoenicica Petrovitz, 1969 Maladera punctatissima (Faldermann, 1836) Maladera rugosa (Blanchard, 1850) Maladera seleuciensis Petrovitz, 1969 Maladera syriaca Petrovitz, 1969 Maladera taurica Petrovitz, 1969 Maladera vignai Sabatinelli, 1977 Maladera wewalkai Petrovitz, 1969 Maladera Maladera alibagensis Ahrens & Fabrizi, 2016 Maladera allonitens Ahrens, Fabrizi & Liu, 2021 Maladera allopruinosa (Ahrens, 1998) Maladera alloservitrita Sreedevi, Speer, Fabrizi & Ahrens, 2018 Maladera amamiana Nomura, 1959 Maladera amboliensis Ahrens & Fabrizi, 2016 Maladera anaimalaiensis Ahrens & Fabrizi, 2016 Maladera analis (Brenske, 1899) Maladera andamana (Brenske, 1899) Maladera anderssoni Fabrizi & Ahrens, 2014 Maladera andrewesi Ahrens & Fabrizi, 2016 Maladera annamensis (Moser, 1915) Maladera antispinosa Ahrens & Fabrizi, 2016 Maladera apicalis Ahrens, Fabrizi & Liu, 2021 Maladera aptera Ahrens, Fabrizi & Liu, 2021 Maladera assamica (Moser, 1915) Maladera aureola (Murayama, 1938) Maladera badullana Fabrizi & Ahrens, 2014 Maladera bagmatiensis Ahrens, 2004 Maladera baii Ahrens, Fabrizi & Liu, 2021 Maladera baishaoensis Ahrens, Fabrizi & Liu, 2021 Maladera bakeri (Moser, 1915) Maladera balphakramensis Ahrens & Fabrizi, 2016 Maladera bandarwelana Fabrizi & Ahrens, 2014 Maladera bansongchana Ahrens, Fabrizi & Liu, 2021 Maladera baoxingensis Ahrens, Fabrizi & Liu, 2021 Maladera barasingha Gupta, Bhunia, Ahrens & Chandra, 2025 Maladera basalis (Moser, 1915) Maladera bawanglingana Ahrens, Fabrizi & Liu, 2021 Maladera bawanglingensis Ahrens, Fabrizi & Liu, 2021 Maladera beibengensis Ahrens, Fabrizi & Liu, 2021 Maladera beidouensis Ahrens, Fabrizi & Liu, 2021 Maladera bengalensis (Brenske, 1899) Maladera bezdeki Bhunia, Gupta, Sarkar & Ahrens, 2023 Maladera bhutanensis (Frey, 1975) Maladera bikouensis Ahrens, Fabrizi & Liu, 2021 Maladera bismarckiana (Brenske, 1899) Maladera bisornata Fabrizi & Ahrens, 2014 Maladera botrytibia Nomura, 1974 Maladera breviata (Brenske, 1899) Maladera breviatella Fabrizi & Ahrens, 2014 Maladera breviclava Ahrens, Fabrizi & Liu, 2021 Maladera brevistylis Ahrens, 2004 Maladera brincki Fabrizi & Ahrens, 2014 Maladera brunnescens (Frey, 1972) Maladera bubengensis Ahrens, Fabrizi & Liu, 2021 Maladera burmeisteri (Brenske, 1899) Maladera calcarata (Brenske, 1899) Maladera calicutensis (Frey, 1972) Maladera cambodjana (Frey, 1972) Maladera cardamomensis Ahrens & Fabrizi, 2016 Maladera celebensis (Moser, 1915) Maladera cervicornis Ranasinghe, Eberle, Benjamin & Ahrens, 2020 Maladera champhaiensis Gupta, Bhunia, Ahrens & Chandra, 2025 Maladera chenzhouana Ahrens, Fabrizi & Liu, 2021 Maladera chinensis (Moser, 1915) Maladera chiruwae Ahrens, 2004 Maladera clavata (Frey, 1972) Maladera coimbatoreensis Ahrens & Fabrizi, 2016 Maladera comosa (Brenske, 1899) Maladera conspicua Ahrens, 2004 Maladera constans Ahrens & Fabrizi, 2016 Maladera constellata Ahrens, Fabrizi & Liu, 2021 Maladera consularis Ahrens & Fabrizi, 2009 Maladera contracta (Brenske, 1899) Maladera costigera (Blanchard, 1850) Maladera coxalis (Moser, 1915) Maladera crenatotibialis Ahrens, Fabrizi & Liu, 2021 Maladera crenolatipes Ahrens, Fabrizi & Liu, 2021 Maladera crinifrons (Brenske, 1899) Maladera curtipes (Moser, 1915) Maladera daanensis Ahrens, Fabrizi & Liu, 2021 Maladera dadongshanica Ahrens, Fabrizi & Liu, 2021 Maladera dahanshana Kobayashi, 2016 Maladera dajuensis Ahrens, Fabrizi & Liu, 2021 Maladera dambullana Ranasinghe, Eberle, Athukorala, Benjamin & Ahrens, 2022 Maladera danfengensis Ahrens, Fabrizi & Liu, 2021 Maladera davaoana (Moser, 1921) Maladera dayaoshanica Ahrens, Fabrizi & Liu, 2021 Maladera declarata Ahrens & Fabrizi, 2016 Maladera decolor Ahrens & Fabrizi, 2016 Maladera deenstana Ranasinghe, Eberle, Athukorala, Benjamin & Ahrens, 2022 Maladera densipilosa Ahrens & Fabrizi, 2016 Maladera detersa (Erichson, 1834) Maladera diaolinensis Ahrens, Fabrizi & Liu, 2021 Maladera dimidiata Ahrens & Fabrizi, 2016 Maladera distincta (Moser, 1915) Maladera diyalumana Fabrizi & Ahrens, 2014 Maladera drescheri (Moser, 1913) Maladera dubia (Arrow, 1916) Maladera duvivieri (Brenske, 1896) Maladera eclogaria (Brenske, 1899) Maladera egregia (Arrow, 1946) Maladera ejimai Miyake & Imasaka, 1987 Maladera eluctabilis (Brenske, 1899) Maladera emeifengensis Ahrens, Fabrizi & Liu, 2021 Maladera engana (Brenske, 1902) Maladera eremita (Brenske, 1899) Maladera erlangshanica Ahrens, Fabrizi & Liu, 2021 Maladera eusericina Ahrens & Fabrizi, 2016 Maladera excisiceps (Frey, 1972) Maladera excisilabrata Ahrens, Fabrizi & Liu, 2021 Maladera exigua (Brenske, 1894) Maladera eximia (Arrow, 1946) Maladera fangana Ahrens, Fabrizi & Liu, 2021 Maladera fasciculata (Moser, 1922) Maladera fastuosa Ahrens & Fabrizi, 2016 Maladera fatigata Ahrens, 2004 Maladera femorata (Brenske, 1899) Maladera fengyangshanica Ahrens, Fabrizi & Liu, 2021 Maladera ferekanarana Ahrens & Fabrizi, 2016 Maladera fereobscurata Ahrens, Fabrizi & Liu, 2021 Maladera ferruginea (Kollar & Redtenbacher, 1848) Maladera festina (Brenske, 1899) Maladera filigraniforceps Ahrens, Fabrizi & Liu, 2021 Maladera fistulosa (Brenske, 1899) Maladera flammea (Brenske, 1899) Maladera flavipennis Ahrens, Fabrizi & Liu, 2021 Maladera flinti Fabrizi & Ahrens, 2014 Maladera floresina (Brenske, 1899) Maladera freyi Ahrens & Fabrizi, 2016 Maladera fuanensis Ahrens, Fabrizi & Liu, 2021 Maladera fulgida (Brenske, 1899) Maladera fuliginosa (Burmeister, 1855) Maladera fumosa (Brenske, 1899) Maladera fuscescens (Moser, 1917) Maladera fuscipes (Moser, 1915) Maladera futschauana (Brenske, 1898) Maladera galdaththana Ranasinghe, Eberle, Benjamin & Ahrens, 2020 Maladera ganglbaueri (Brenske, 1899) Maladera gansuensis (Miyake & Yamaya, 2001) Maladera garoana Ahrens & Fabrizi, 2016 Maladera geniculata Ahrens & Fabrizi, 2016 Maladera gibbiventris (Brenske, 1898) Maladera gopaldharae Ahrens, 2004 Maladera gorkhae Ahrens, 2004 Maladera granigera Ahrens & Fabrizi, 2016 Maladera granuligera (Blanchard, 1850) Maladera guangzhaishanica Ahrens, Fabrizi & Liu, 2021 Maladera guanxianensis Ahrens, Fabrizi & Liu, 2021 Maladera guanxiensis Ahrens, Fabrizi & Liu, 2021 Maladera guomenshanica Ahrens, Fabrizi & Liu, 2021 Maladera gusakovi Ahrens, Fabrizi & Liu, 2021 Maladera guttula (Sharp, 1876) Maladera haba Ahrens, Fabrizi & Liu, 2021 Maladera habashanensis Ahrens, Fabrizi & Liu, 2021 Maladera haldwaniensis Ahrens, 2004 Maladera hampsoni Ahrens & Fabrizi, 2016 Maladera haniel Ranasinghe, Eberle, Athukorala, Benjamin & Ahrens, 2022 Maladera hansmalickyi Ahrens, Fabrizi & Liu, 2021 Maladera harmonica (Brenske, 1898) Maladera hastata Fabrizi & Ahrens, 2014 Maladera hauseri (Brenske, 1899) Maladera hayashii Hirasawa, 1991 Maladera heveli Fabrizi & Ahrens, 2014 Maladera hiemalis Kobayashi, 2014 Maladera hiyarensis Ranasinghe, Eberle, Athukorala, Benjamin & Ahrens, 2022 Maladera hmong Ahrens, 2004 Maladera holosericea (Scopoli, 1772) Maladera holzschuhi Ahrens, 2004 Maladera hongyuanensis Ahrens, Fabrizi & Liu, 2021 Maladera horii (Kobayashi, 2010) Maladera hortonensis Fabrizi & Ahrens, 2014 Maladera houzhenziensis Ahrens, Fabrizi & Liu, 2021 Maladera howdeni Ahrens, 2003 Maladera huanianensis Ahrens, Fabrizi & Liu, 2021 Maladera hubeiensis Ahrens, Fabrizi & Liu, 2021 Maladera hui Ahrens, Fabrizi & Liu, 2021 Maladera hunanensis Ahrens, Fabrizi & Liu, 2021 Maladera hunliensis Ahrens & Fabrizi, 2016 Maladera hunuguensis Ahrens, Fabrizi & Liu, 2021 Maladera hutiaoensis Ahrens, Fabrizi & Liu, 2021 Maladera iliganica (Moser, 1917) Maladera imasakai Miyake & Yamaya, 1995 Maladera imbella Reitter, 1898 Maladera impressithorax Nomura, 1973 Maladera impubis Ahrens, 2004 Maladera inadai (Kobayashi, 2010) Maladera inaequabilis (Brenske, 1899) Maladera indica (Blanchard, 1850) Maladera inermis (Brenske, 1899) Maladera inimica (Brenske, 1899) Maladera initialis Ahrens & Fabrizi, 2016 Maladera inornata (Brenske, 1899) Maladera insularis (Brenske, 1899) Maladera invenusta (Moser, 1918) Maladera iridicauda (Fairmaire, 1893) Maladera irididorsis Ahrens, 2004 Maladera isarogensis (Moser, 1922) Maladera iuga Fabrizi & Ahrens, 2014 Maladera jaintiaensis Ahrens & Fabrizi, 2016 Maladera japonica (Motschulsky, 1860) Maladera jaroslavi Ahrens, Fabrizi & Liu, 2021 Maladera jatuai Ahrens, Fabrizi & Liu, 2021 Maladera jiangi Ahrens, Fabrizi & Liu, 2021 Maladera jingdongensis Ahrens, Fabrizi & Liu, 2021 Maladera jinggangshanica Ahrens, Fabrizi & Liu, 2021 Maladera jinghongensis Ahrens, Fabrizi & Liu, 2021 Maladera jiucailingensis Ahrens, Fabrizi & Liu, 2021 Maladera jizuana Ahrens, Fabrizi & Liu, 2021 Maladera judithae Sreedevi & Ahrens, 2025 Maladera kaimurensis Chandra, Ahrens, Bhunia, Sreedevi & Gupta, 2021 Maladera kalawensis Ahrens, Fabrizi & Liu, 2021 Maladera kallarensis Ahrens & Fabrizi, 2016 Maladera kamiyai (Sawada, 1937)