Heteronyx is a genus of beetles in the family Melolonthidae found in Australia and Melanesia.
Taxonomic history The genus has been challenging for scientists, because many species are very similar in appearance. Thomas Blackburn described most of the Australian species, using the key he devised to separate the species. Arthur Mills Lea spend years working on a revision of the genus, but eventually gave up. It was not until 2000 that E. B. Britton published a revision of the genus, in which he synonomised 111 of the total of 353 described Australian species. Britton states that the aim of his revision was to establish synonymies and to make identifiable the valid species so far described. This would make a future full revision of the genus possible and would also make it possible to describe the numerous species that are yet undescribed.
Species
Heteronyx acutifrons Blackburn, 1889 Heteronyx additus Blackburn, 1909 Heteronyx advena Blackburn, 1890 Heteronyx aequaliceps Blackburn, 1890 Heteronyx aequalis Blackburn, 1888 Heteronyx affinis Blackburn, 1909 Heteronyx agrestis Burmeister, 1855 Heteronyx agricola Blackburn, 1910 Heteronyx alienus Blackburn, 1892 Heteronyx alpicola Blackburn, 1892 Heteronyx ambiguus Blackburn, 1910 Heteronyx amboinensis Moser, 1920 Heteronyx apertus Blackburn, 1910 Heteronyx aphodioides Blanchard, 1850 Heteronyx apicispinosa Heller, 1914 Heteronyx arcanus Blackburn, 1892 Heteronyx aridus Blackburn, 1889 Heteronyx aruanus Moser, 1920 Heteronyx asperatus Moser, 1920 Heteronyx aspericollis Blackburn, 1889 Heteronyx asperifrons Blackburn, 1909 Heteronyx augustae Blackburn, 1889 Heteronyx australis Guérin-Méneville, 1831 Heteronyx badius MacLeay, 1888 Heteronyx beltanae Blackburn, 1888 Heteronyx bidentatus Blackburn, 1889 Heteronyx blandus Blackburn, 1909 Heteronyx borealis Blackburn, 1889 Heteronyx bovilli Blackburn, 1890 Heteronyx brevior Fairmaire, 1883 Heteronyx caledoniae Fauvel, 1903 Heteronyx calidus Blackburn, 1910 Heteronyx callabonnae Blackburn, 1909 Heteronyx capillatus MacLeay, 1888 Heteronyx castanescens Moser, 1924 Heteronyx castaneus Blanchard, 1850 Heteronyx cervina (Boisduval, 1835) Heteronyx chlorotica (Gyllenhal, 1817) Heteronyx cliens Blackburn, 1910 Heteronyx collaris Blackburn, 1890 Heteronyx colossus Arrow, 1915 Heteronyx comans Blackburn, 1909 Heteronyx concolor MacLeay, 1871 Heteronyx confertus Blackburn, 1910 Heteronyx constans Blackburn, 1889 Heteronyx corpulentus MacLeay, 1888 Heteronyx costulatus Blackburn, 1910 Heteronyx crassus Blackburn, 1889 Heteronyx cribriceps Blackburn, 1892 Heteronyx crinitus Blackburn, 1910 Heteronyx cunnamullae Blackburn, 1910 Heteronyx cygneus Blackburn, 1889 Heteronyx darlingensis Blackburn, 1888 Heteronyx declaratus Blackburn, 1909 Heteronyx dentipes Blackburn, 1889 Heteronyx dimidiatus (Erichson, 1842) Heteronyx disjectus Blackburn, 1910 Heteronyx diversiceps Blackburn, 1890 Heteronyx doctus Blackburn, 1890 Heteronyx doddi Blackburn, 1908 Heteronyx electus Blackburn, 1889 Heteronyx elongatus Blanchard, 1850 Heteronyx elytrurus Blackburn, 1909 Heteronyx excisa Heller, 1912 Heteronyx excisus Blackburn, 1890 Heteronyx exectus Blackburn, 1909 Heteronyx fervidus Blackburn, 1892 Heteronyx fimbriatus Moser, 1924 Heteronyx flavus Blackburn, 1890 Heteronyx fortis Blackburn, 1888 Heteronyx fraserensis Blackburn, 1892 Heteronyx frenchi Blackburn, 1909 Heteronyx froggatti MacLeay, 1888 Heteronyx frontalis Blackburn, 1888 Heteronyx fulvohirtus Blackburn, 1888 Heteronyx fumata (Erichson, 1842) Heteronyx furvus Blackburn, 1892 Heteronyx glabratus (Erichson, 1842) Heteronyx grandis Blackburn, 1900 Heteronyx granulatus Blackburn, 1910 Heteronyx granulifer Blackburn, 1889 Heteronyx granum Burmeister, 1855 Heteronyx helmsi Blackburn, 1892 Heteronyx hirsutosetosus Moser, 1920 Heteronyx hirsutus Blackburn, 1910 Heteronyx hirtuosus Blackburn, 1890 Heteronyx hispidulus Blackburn, 1908 Heteronyx horridus Blackburn, 1888 Heteronyx humilis Blackburn, 1910 Heteronyx imitator Blackburn, 1909 Heteronyx impar Blackburn, 1910 Heteronyx incognitus Blackburn, 1892 Heteronyx incola Blackburn, 1889 Heteronyx infirmus Blackburn, 1910 Heteronyx infuscatus MacLeay, 1871 Heteronyx inornatus (Blackburn, 1900) Heteronyx insignis Blackburn, 1888 Heteronyx insularis Fairmaire, 1883 Heteronyx interioris Blackburn, 1909 Heteronyx intrusus Blackburn, 1910 Heteronyx iridiventris Blackburn, 1890 Heteronyx irrasus Lea, 1926 Heteronyx jejunus Blackburn, 1888 Heteronyx johannis Blackburn, 1912 Heteronyx labratus Moser, 1926 Heteronyx laeviceps Blackburn, 1888 Heteronyx laminatus Blackburn, 1890 Heteronyx lateritius Blackburn, 1889 Heteronyx laticeps Burmeister, 1855 Heteronyx lilliputanus Blackburn, 1890 Heteronyx lindi Blackburn, 1888 Heteronyx lividus Blackburn, 1889 Heteronyx lobatus Blackburn, 1888 Heteronyx luteolus Blackburn, 1909 Heteronyx maculatus Blackburn, 1888 Heteronyx magnus Moser, 1924 Heteronyx major Blackburn, 1910 Heteronyx maluensis Brenske, 1889 Heteronyx marcidus Blackburn, 1892 Heteronyx marginatus Blackburn, 1890 Heteronyx merus Blackburn, 1892 Heteronyx metropolitanus Blackburn, 1909 Heteronyx mimus Blackburn, 1890 Heteronyx minimus Lea, 1926 Heteronyx minutus Blackburn, 1910 Heteronyx moluccanus Moser, 1920 Heteronyx montanus Blackburn, 1890 Heteronyx mulwalensis Blackburn, 1889 Heteronyx mundus Blackburn, 1909 Heteronyx nasutus Blackburn, 1889 Heteronyx neglectus Blackburn, 1910 Heteronyx nigellus (Erichson, 1842) Heteronyx nigrescens Blackburn, 1909 Heteronyx nigricans Burmeister, 1855 Heteronyx nigrita Blanchard, 1850 Heteronyx nigritulus Moser, 1926 Heteronyx normalis Blackburn, 1889 Heteronyx novitius Blackburn, 1910 Heteronyx obesa Burmeister, 1855 Heteronyx oblongus Blanchard, 1850 Heteronyx orbus Blackburn, 1909 Heteronyx ovatus Blanchard, 1850 Heteronyx pallidulus MacLeay, 1871 Heteronyx paniei Paulian, 1991 Heteronyx papuanus Moser, 1920 Heteronyx parvulus MacLeay, 1888 Heteronyx pauxillus Blackburn, 1910 Heteronyx pellucidus Burmeister, 1855 Heteronyx peregrinus Blackburn, 1890 Heteronyx perkinsi Blackburn, 1909 Heteronyx piceoniger MacLeay, 1888 Heteronyx piceus Blanchard, 1850 Heteronyx pilosellus Blanchard, 1850 Heteronyx pilosus Blanchard, 1850 Heteronyx placidus Blackburn, 1910 Heteronyx planiceps Blackburn, 1910 Heteronyx posticalis Blackburn, 1890 Heteronyx praecox (Erichson, 1842) Heteronyx proditor Blackburn, 1892 Heteronyx proprius Blackburn, 1910 Heteronyx prosper Blackburn, 1909 Heteronyx protervus Blackburn, 1892 Heteronyx proxima Burmeister, 1855 Heteronyx pubescens (Erichson, 1842) Heteronyx pumilus Sharp, 1877 Heteronyx puncticollis Blackburn, 1890 Heteronyx punctipennis Blackburn, 1889 Heteronyx pustulosus Blackburn, 1890 Heteronyx pygidialis Blackburn, 1888 Heteronyx pygmaeus Blackburn, 1910 Heteronyx quadraticollis Blackburn, 1890 Heteronyx queenslandicus Blackburn, 1909 Heteronyx randalli Blackburn, 1890 Heteronyx rectangulus Blackburn, 1910 Heteronyx relictus Blackburn, 1909 Heteronyx rhinastus Blackburn, 1890 Heteronyx rhinoceros Blackburn, 1892 Heteronyx rothei Blackburn, 1890 Heteronyx rotundiceps Blanchard, 1850 Heteronyx rubescens Blanchard, 1850 Heteronyx ruficollis MacLeay, 1871 Heteronyx rufomarginatus Blanchard, 1850 Heteronyx rufopiceus MacLeay, 1888 Heteronyx rugosipennis MacLeay, 1871 Heteronyx rusticus Blackburn, 1890 Heteronyx salebrosus Blackburn, 1909 Heteronyx satelles Blackburn, 1888 Heteronyx scalptus Blackburn, 1890 Heteronyx sequens Blackburn, 1894 Heteronyx setifer Blackburn, 1890 Heteronyx sexualis Blackburn, 1909 Heteronyx siccus Blackburn, 1892 Heteronyx simius Blackburn, 1890 Heteronyx simplicicollis Blackburn, 1909 Heteronyx simulator Blackburn, 1888 Heteronyx sloanei Blackburn, 1889 Heteronyx solidus Blackburn, 1888 Heteronyx spadiceus Burmeister, 1855 Heteronyx spretus Blackburn, 1888 Heteronyx squalidus Blackburn, 1908 Heteronyx striatus Blackburn, 1909 Heteronyx suavis Blackburn, 1909 Heteronyx subferruginea Burmeister, 1855 Heteronyx subfuscus MacLeay, 1888 Heteronyx subglaber MacLeay, 1888 Heteronyx substriatus MacLeay, 1871 Heteronyx sulcifrons Blackburn, 1909 Heteronyx sydneyanus Blackburn, 1890 Heteronyx tarsalis Blackburn, 1909 Heteronyx tasmanicus Blackburn, 1909 Heteronyx tenebrosus Blackburn, 1909 Heteronyx tepperi Blackburn, 1888 Heteronyx terrena Blackburn, 1892 Heteronyx tindalei Britton, 1992 Heteronyx torvus Blackburn, 1888 Heteronyx transversicollis MacLeay, 1888 Heteronyx tridentatus Lea, 1924 Heteronyx tristis Blackburn, 1888 Heteronyx umbilicatus Fauvel, 1903 Heteronyx unicolor Blanchard, 1850 Heteronyx unicus Blackburn, 1900 Heteronyx variegatus Blackburn, 1888 Heteronyx vicinus Blackburn, 1910 Heteronyx victoris Blackburn, 1888 Heteronyx viduus Blackburn, 1910 Heteronyx zalotus Blackburn, 1910
Species with unclear status Heteronyx planatus Burmeister, 1855 (described as Heteronyx planata from Western Australia)
References