Protopteraspididae is an extinct family of pteraspidid heterostracan agnathans. Fossils of the various genera are found in early Devonian-aged marine strata. Protopteraspidids were once thought to represent a taxon of basal pteraspidids (hence "proto") but recent evaluations demonstrate that Protopteraspididae is a paraphyletic group of various transitional forms representing a gradual transition between the more advanced Pteraspoidei (comprising Pteraspididae, Gigantaspis and Protaspididae),Pernegre, Vincent N., and David K. Elliott. "Phylogeny of the Pteraspidiformes (Heterostraci), Silurian–Devonian jawless vertebrates." Zoologica Scripta 37.4 (2008): 391-403. and the anchipteraspidids and the Psammosteids.

Genera Protopteraspids are found in Lower Devonian marine strata of the Western United States, Northern Canada, England, France, Belgium, Svalbard, Norway, and Podolia, Ukraine.Blieck, Alain. "Les hétérostracés Ptéraspidiformes, agnathes du Silurien-Dévonien du continent Nord-Atlantique et des blocs avoisinants: Révision systématique, phylogénie, biostratigraphie, biogéographie." (1984).

Protopteraspis This genus is known from several species from Lower Devonian England, France and Belgium

Doryaspis This is a genus of aberrant pteraspidids with lower lips elongated into long, dagger-like organs of unknown function. The various species are primarily restricted to the Lower Devonian strata of Svalbard.

Escharaspis Miltaspis Stegobranchiaspis Unarkaspis "Trygonaspis" "Trygonaspis" is a nomen nudum given to a well-preserved complete armor that strongly resembles Protopteraspis in form, but with a long, recurved dorsal spine, and orbital plates that have bookshelf-like extensions. The only known specimen was found in Northern Canada.

References

Palaeos Vertebrates entry on order Pteraspidiformes