The Amazon biome () contains the Amazon rainforest, an area of tropical rainforest, and other ecoregions that cover most of the Amazon basin and some adjacent areas to the north and east. The biome contains blackwater and whitewater flooded forest, lowland and montane terra firma forest, bamboo and palm forest, savanna, sandy heath and alpine tundra. Some areas of the biome are threatened by deforestation for timber and to make way for pasture or soybean plantations.
Location
The Amazon biome has an area of . The biome roughly corresponds to the Amazon basin, but excludes areas of the Andes to the west and cerrado (savannah) to the south, and includes lands to the northeast extending to the Atlantic ocean with similar vegetation to the Amazon basin. J. J. Morrone (2006) defines the Amazonian subregion in this broader sense, divided into the biogeographical provinces of Guyana, Humid Guyana, Napo, Imeri, Roraima, Amapá, Várzea, Ucayali, Madeira, Tapajós-Xingu, Pará, Yungas and Pantanal. The World Wildlife Fund takes a similar scope, where the Amazon biome includes the Guiana Shield rain forests in the north and the Chiquitano dry forests of Bolivia.
The biome covers parts of Brazil, Bolivia, Peru, Ecuador, Colombia, Venezuela, Guyana, Suriname and French Guiana. In Brazil the biome covers more than and covers all or parts of the states of Acre, Amazonas, Roraima, Rondônia, Pará, Amapá, Maranhão, Tocantins and Mato Grosso. The Amazon biome covers 49.29% of Brazil. 16% of the biome is in Peru. As of 2015 about 23.4% of Peru's Amazon biome was protected, but of this less than half was fully protected.
Physical environment Terrain
Much of the terrain of the Amazon biome, particularly around the rivers, is lowland plains. The Guiana Shield is an area of highlands along the border between Brazil and Venezuela and Guyana. The southern Amazonian highlands cross parts of Rondonia and Mato Grosso and the southern parts of Amazonas and Para.
The Amazon basin is crossed by ridges or "paleoarches" that connect the Guiana and Central Brazil shields and divide it into geological sub-basins. They are the Iquitos or Jutai Arch in Peru and Acre, the Carauari Arch across the Rio Negro and Solimões, the Purus Arch to the west of Manaus, the Monte Alegre Arch to the west of the Tapajós and the Gurupa Arch to the west of Marajó. Under the Paleoarch model, paleobasins between the arches form centers for biological diversification. Thus the Iquitos arch is considered the main reason for the different species of frogs and rodents and different forest types on either side of the ridge.
Soil
The soil is generally very poor in nutrients, and areas that have been deforested are often unsuitable for agriculture or pasture. A large part of the phosphorus necessary for life is blown by the wind from Africa; as diatomeous dust from the Bodélé Depression and as smoke due to biomass burning in Africa. There are wide regional variations in soil types. Thus 20% of the Rio Negro basin is covered by podzols and 55% by acrisols and ferralsols, with the remainder covered by alluvial and litholic soils and scattered areas of hydromorphic plinthosols. In the biome as a whole podzols cover just , or 2.7% of the area.
Climate In Brazil the average temperature of the biome is and average rainfall is , but there are wide variations from one region to another. The biome as a whole has annual rainfall from , about half of which is carried by winds from the Atlantic, and the other half from evapotranspiration within the biome. There are wide variations in total rainfall and distribution of rainfall throughout the year.
Water
The Amazon watershed covers about . The Amazon River accounts for 15–16% of the total water discharged by rivers into the oceans of the world. Rivers may be blackwater, whitewater or clearwater. Thus the Rio Negro ("Black River") has clear, jet-black water caused by decomposition of organic matter in swamps along its margins, combined with low levels of silt. The Rio Branco ("White River") and the Amazon itself have yellowish waters loaded with silt. The Tahuayo River in the Tamshiyacu Tahuayo Regional Conservation Area of Peru is classed as a blackwater river, but often has similar chemistry to the whitewater rivers of the region since it is in the Amazon River floodplain, and receives water from the Amazon.
The Amazon and its major tributaries such as the Xingu, Tapajós, Madeira, Purus and Rio Negro form barriers to the geodispersal of plants, animals and even insects. Thus the white-fronted capuchin (Cebus albifrons) and hairy saki (Pithecia hirsuta) are found west of the Tapajós, while the white-nosed saki (Chiropotes albinasus) is only found east of the river. The World Wildlife Fund divides the biome into ecoregions, often defined as the regions lying between major tributaries of the Amazon.
Ecosystems
Most of the interior of the Amazon basin is covered by rainforest. The dense tropical Amazon rainforest is the largest tropical rainforest in the world. It covers between of the Amazon biome. The somewhat vague numbers are because the rainforest merges into similar biomes across its boundaries. The rainforest is Tropical Broadleaf Forest, so-called because most of the trees have broad leaves. The basin also holds flooded riparian forest or várzea, seasonal forest and savanna. Seasonal forest covers much of the southeast border, with marked dry seasons when there are frequent fires. The Amazon biome contains areas of other types of vegetation including grasslands, swamps, bamboos, and palm forests.
There are 53 major ecosystems and more than 600 types of land and freshwater habitat. Of the ecosystems, 34 are forest areas covering 78% of the biome, 6 are Andean covering 1.5%, 5 are floodplains covering 5.83%, 5 are savanna covering 12.75% and two are tropical steppes covering 1.89%. The Brazilian Amazon holds 30 of the 53 ecosystems, of which 19 are forests with 77.5% of the area. The borders of the biome hold ecotones where it blends into other biomes such as the cerrado.
Within and across the ecosystems of the biome there is huge biological diversity. One source says there are an estimated 60,000 plant species, of which 30,000 are endemic. Another says there are 30,000 to 50,000 plant species. The Brazilian Ministry of the Environment as of January 2013 listed 2,500 species of trees and 30,000 species of plants. There are 1,400 species of fish, 163 amphibians, 387 reptiles and more than 500 mammals including 90 primates. 87% of the amphibians, 62% of reptiles, 20% of birds and 25% of mammals are endemic to the biome. 109 species of lizards and amphisbaena reptiles are known to be present, and 138 species of snakes. Of the 1,300 bird species 20% are endemic and 8.4% are endangered.
A relatively small area may contain a variety of ecoregions. Thus the Pico da Neblina National Park in the north of Amazonas, Brazil contains campinarana, dense rainforest and contact between campinarana and rainforest. The vegetation formations include terra firme forests and igapós. Submontane forests climb the first steps of the Guiana Plateau to about , followed by montane forests. Lichens and bromeliads are found up to . There is also alpine tundra in the tabular plateaus.
Moist forest
The Amazon basin contains several large areas of moist forest, collectively called the Amazon rainforest. These are the Caquetá, Japurá–Solimões–Negro, Juruá–Purus, Madeira–Tapajós, Napo, Purus–Madeira, Solimões–Japurá, Southwest Amazon, Tapajós–Xingu, Tocantins–Araguaia–Maranhão, Uatuma–Trombetas, Ucayali, and Xingu–Tocantins–Araguaia moist forests. Each has distinctive vegetation. In the broad definition favored by the World Wide Fund for Nature (WWF) the Amazon biome would also include the Tepuis, table-top mountains with unique vegetation, the Negro-Branco, Guianan Highlands, Guianan piedmont and lowland and Guianan moist forests and the Guianan savanna.
In the southwest Amazon at least of forest are dominated by bamboos. These occur in areas where there has been recent tectonic uplift combined with fast mechanical erosion and poor drainage. In each patch of bamboos all the plants flower every 27–28 years, producing huge numbers of seeds, before dying. The Chandless State Park is in the centre of the bamboo forest region of the south western Amazon biome, and has three endemic species of the Guadua genus of bamboo. Vegetation is classified as forest with palms, forest with bamboo, alluvial forest with bamboo and dense forest.
Flooded forest
Before Ghillean Prance's 1979 study, the local people and many scientists called the permanently waterlogged swamp on the lower Amazon "igapó" and all types of periodically flooded land "várzea". A more recent definition, from Prance, is: Type of flooding Type of water Name Alternative namesSeasonalWhitewater Seasonal várzea Blackwater Seasonal igapó TidalSalt water Mangrove Fresh water back-upTidal várzea Irregular rainfall Floodplain forest Riparian forest along low order riversPermanentWhitewater Permanent swamp forest várzea swamp forest, várzea chavascalBlackwater and clearwater Permanent igapó Long-term inundated forests Major flooded areas include the Iquitos, Purus, Monte Alegre, Gurupa and Marajó várzeas. The Majaró várzea is at the mouth of the Amazon and is affected by both freshwater and tidal flows.
Campina and campinarana
Campina is open forest on sandy soil where sunlight can reach the ground. More than half the species of orchid in the Amazon lowlands are found in this type of forest. The terms campina and campinarana both describe white sand savannas that are very poor in nutrients. They may be flooded periodically or seasonally, in which case the roots suffer from lack of aeration. The vegetation is stunted. For some, campina is distinguished as being completely treeless. The term "campinarana" is used for the transition between campina and the rainforest.
Campinarana is found in isolated areas of the Rio Negro and Rio Branco basins in the north of Brazil. Patches of campinarana, which may cover several thousand square kilometres, are found in the transitional region from the Guyana Shield to the Amazon basin. Campinarana is typically found on leached white sands around circular swampy depressions in lowland tropical moist forest. The soil is low in nutrients, with highly acidic humus. Campinarana includes savannah, scrub and forests. The savannah is mainly composed of grasses and lichens, found in the wet plains beside lakes and rivers. The scrub has bare sand, herbaceous plants, shrubs and trees less than high.
The nutrient-poor Japurá-Solimões-Negro moist forest is similar to the Rio Negro Campinarana.
Restinga
The Northeastern Brazil restingas are coastal dune habitats that extend along the coast of northeastern Brazil, interspersed with lagoons, mangroves and patches of caatinga savanna. The land behind the dunes may include dwarf palms, bromeliads, ferns, shrubs, grasses and scrub trees. The more exposed areas mainly hold medium-tall grasses and scrub trees, while sheltered areas hold patches of cactus and low dry thicket. Fauna include marmosets and jaguarundis, proboscis bats, lesser sac-winged bats, bulldog bats, and Davy's naked-backed bats, wood stork, roseate spoonbill, white-necked heron, great egret, cattle egret, black-crowned night heron, and Neotropic cormorant. The ecoregion has various endemic species. The Lençóis Maranhenses National Park protects a large area of this habitat.
Conservation
Of the 1,149 federal and state protected areas in Brazil in 2014, 247 covering were in the Amazon biome and accounted for 73% of the total area protected at the federal and state levels in Brazil. Federal units covered and state units covered .