The Mishash Formation is a Late Cretaceous (Campanian-Maastrichtian) geologic formation in Israel. The formation is correlated with the Duwi Formation of Egypt and the Amman and Ruseifa Formations of Jordan.Mishash Formation at Fossilworks.org Mosasaur fossils assigned to Prognathodon currii and pterosaur fossils have been recovered from the formation.
Stratigraphy The lower boundary of the Mishash Formation lies atop the top of the underlying Menuha Formation, while the upper boundary is covered by the overlying Ghareb Formation. The Mishash Formation is divided into two members known as the Chert Member and Phosphate Member, with the latter overlying the former. Massive chert beds comprise the Chert Member, while the Phosphate Member is made up of cherts, phosphorites, porcellanites and organic-rich carbonates. The Phosphate Member is further divided into three units; the Phosphatic Carbonate Unit lies at the bottom of the member, followed by the Porcelanite Unit in the middle, and finally the Phosphorite Unit at the top.
Fossil content
Reptiles Reptiles of the Mishash FormationGenusSpeciesLocationStratigraphic positionMaterialNotesImageArambourgianiaA. sp.Nahal Qazra & OronPhosphate Member (Phosphorite & Phosphatic Carbonate units)Endocranium & bone castsAn azhdarchid pterosaur, originally reported as Titanopteryx.LiodonL. asiaticumNear JerusalemNumerous bonesA dubious mosasaur, remains now assigned to Mosasaurinae indet. & Mosasaurini incertae sedisMosasaurinae indet.IndeterminateNortheastern JerusalemCaudal vertebraeMosasaur remains originally assigned to Liodon asiaticumMosasauriniIncertae sedisNortheastern Jerusalem30 bonesA mosasaur related to Mosasaurus and Plotosaurus, remains originally assigned to Liodon asiaticumPrognathodonP. curriiOronPhosphate MemberSkull & 8 vertebraeA large mosasaurReptilia indet.IndeterminateNahal QazraPhosphate Member (Phosphatic Carbonate Unit)EndocraniumRemains of an indeterminate reptile.TitanopteryxT. sp.Nahal Qazra & OronPhosphate Member (Phosphorite & Phosphatic Carbonate units)Endocranium & bone castsJunior synonym of Arambourgiania.
Bony fish Boney Fish of the Mishash FormationGenusSpeciesLocationStratigraphic positionMaterialNotesImageEnchodusE. faujasiNahal Qazra & OronPhosphate Member (Phosphorite & Phosphatic Carbonate units)Endocranial castsAn aulopiform.E. sp.Nahal Qazra, Nahal Zin, Nahal Ashosh & OronPhosphate Member (Phosphorite, Porcelanite & Phosphatic Carbonate units)Endocranial casts & bonesAn aulopiform.MicropycnodonM. sp.OronPhosphate Member (Phosphorite Unit)3 endocranial castsA pycnodont.PolymixiidaeIndeterminateOronPhosphate Member (Phosphorite Unit)3 complete & a fragmentary endocranial castA beardfish.PrionolepisP. sp.Nahal AshoshPhosphate Member (Phosphorite Unit)Endocranial castAn aulopiform.StratodusS. sp.Nahal Qazra, Nahal Zin, Nahal Ashosh & OronPhosphate Member (Phosphorite & Phosphatic Carbonate units)Endocranial casts & bonesAn aulopiform.
Cartilaginous fish Cartilaginous Fish of the Mishash FormationGenusSpeciesLocationStratigraphic positionMaterialNotesImageAnomotodonA. plicatusOronPhosphate Member (Phosphorite Unit)TeethA goblin sharkA. sp.Nahal ZinimPhosphate Member (Phosphorite Unit)TeethA goblin sharkCentroscymnuscf. C. sp.OronPhosphate Member (Phosphorite Unit)TeethA sleeper sharkCentrosqualusC. sp.OronPhosphate Member (Phosphorite Unit)TeethA dogfish sharkChiloscylliumC. sp.OronPhosphate Member (Phosphatic Carbonate Unit)TeethA carpet sharkCretalamnaC. appendiculataOronPhosphate Member (Phosphorite Unit)TeethA megatooth sharkC. caraibaeaOron & Nahal ZinimPhosphate Member (Phosphorite Unit)TeethSpecies reassigned to SerratolamnaC. maroccanaOron & Nahal ZinimPhosphate Member (Phosphorite Unit)TeethA megatooth sharkC. sp.OronPhosphate Member (Phosphatic Carbonate Unit)TeethA megatooth sharkCretascymnusC. sp.OronPhosphate Member (Phosphorite Unit)TeethA sleeper sharkCretodusC. sp.OronPhosphate Member (Phosphatic Carbonate Unit)TeethA pseudoscapanorhynchid shark, small-sized member of the genusEchinorhinidae indet.IndeterminateOronPhosphate Member (Phosphorite Unit)TeethAn echinorhinid sharkGanopristisG. leptodonOronPhosphate Member (Phosphorite Unit)TeethGenus now deemed a junior synonym of SclerorhynchusG. sp.OronPhosphate Member (Phosphatic Carbonate Unit)TeethHeterodontusH. sp.OronPhosphate Member (Phosphorite Unit)TeethA bullhead sharkHexanchusH. microdonOronPhosphate Member (Phosphorite & Phosphatic Carbonate units)Numerous teethA cow sharkParaorthacodusP. aff. nerviensisOronPhosphate Member (Phosphorite Unit)ToothA paraorthacodontid sharkPristiophorusP. sp.OronPhosphate Member (Phosphorite Unit)TeethA sawsharkPseudocoraxP. aff. affinisOronPhosphate Member (Phosphorite Unit)Many teethA pseudocoracid sharkPteroscylliumP. sp. 1OronPhosphate Member (Phosphorite Unit)TeethA catsharkP. sp. 2OronPhosphate Member (Phosphatic Carbonate Unit)TeethA catsharkRhinobatoidei n. gen.OronPhosphate Member (Phosphatic Carbonate Unit)TeethA guitarfishRhinobatosR. sp. 1OronPhosphate Member (Phosphorite Unit)TeethA guitarfishR. sp. 2OronPhosphate Member (Phosphorite Unit)TeethA guitarfishR. sp. 3OronPhosphate Member (Phosphatic Carbonate Unit)TeethA guitarfishRhombodusR. binkhorstiNahal ZinimPhosphate Member (Phosphorite Unit)TeethA myliobatiform rayScapanorhynchusS. cf. raphiodonOronPhosphate Member (Phosphorite Unit)TeethA goblin sharkSclerorhynchusS. leptodonOronPhosphate Member (Phosphorite & Phosphatic Carbonate units)TeethA sawskate originally reported as Ganopristis leptodonS. sp.OronPhosphate Member (Phosphatic Carbonate Unit)TeethA sawskate, some material originally reported as Ganopristis sp., also found in the Menuha FormationScyliorhinusS. sp.OronPhosphate Member (Phosphorite & Phosphatic Carbonate units)TeethA catsharkSerratolamnaS. caraibaeaOron & Nahal ZinimPhosphate Member (Phosphorite Unit)TeethA serratolamnid sharkSqualicoraxS. bassaniiOronPhosphate Member (Phosphorite Unit)TeethAn anacoracid sharkS. pristodontusOron & Nahal ZinimPhosphate Member (Phosphorite & Phosphatic Carbonate units)TeethAn anacoracid sharkS. sp.Oron & Nahal ZinimPhosphate Member (Phosphorite & Phosphatic Carbonate units)TeethAn anacoracid sharkSqualusS. sp.OronPhosphate Member (Phosphorite Unit)TeethA dogfish sharkSquatinaS. hasseiOronPhosphate Member (Phosphorite Unit)TeethAn angelsharkTriakidaeTriakid 1OronPhosphate Member (Phosphorite Unit)TeethA houndsharkTriakid 2OronPhosphate Member (Phosphorite Unit)TeethA houndshark
See also List of pterosaur-bearing stratigraphic units
References
Further reading
Y. Edelman-Furstenberg. 2008. Macrobenthic community structure in a high-productivity region: Upper Campanian Mishash Formation (Israel). Palaeogeography, Palaeoclimatology, Palaeoecology 261:58-77 P. Christiansen and N. Bonde. 2002. A new species of gigantic mosasaur from the Late Cretaceous of Israel. Journal of Vertebrate Paleontology 22(3):629-644 Z. Lewy and H. Cappetta. 1989. Senonian Elasmobranch teeth from Israel, biostratigraphic an paleoenvironmental implications. Neues Jahrbuch für Geologie und Paläontologie - Monatshefte 1989(4):212-222