Scythris is a genus of gelechioid moths. It is the type genus of the flower moth family, which is sometimes included as a subfamily in the Xyloryctidae, or together with these merged into the Oecophoridae.Pitkin & Jenkins (2004), AEBR (2008), ToL (2008), FE (2009), and see references in Savela (2008) The genus was erected by Jacob Hübner in 1825.
It is the largest genus of flower moths, and as such might not be fully monophyletic with regard to some very small or monotypic genera placed in the same family. In addition, new species of Scythris continue to be discovered and described.FE (2009), and see references in Savela (2008)
Selected species
Species groups The species of Scythris have been divided among several groups, which may or may not be monophyletic; those that are may – provided they are closely enough related to the type species S. limbella – be considered subgenera. There is a considerable number of species whose exact relationships are hitherto elusive. Some have been placed in a "monospecific species group" of their own, as they are too distinct from the other groups to be included there, yet at the same time have characteristic autapomorphies. Species groups include:AEBR (2008), FE (2009), and see references in Savela (2008) aenea group Scythris aenea Passerin d'Entrèves 1984 aerariella group Scythris adustella Jäckh, 1978 Scythris aerariella (Herrich-Schäffer, 1855) Scythris anomaloptera (Staudinger, 1880) Scythris binotiferella (Ragonot, 1880) Scythris brummanae Passerin d'Entrèves & Roggero, 2012 Scythris carboniella Jäckh, 1978 Scythris corsa Passerin d'Entrèves, 1986 Scythris dorycniella (Milliére, 1861) Scythris flaviventrella (Herrich-Schäffer, 1855) Scythris hungaricella Rebel, 1917 Scythris imperiella Jäckh, 1978 Scythris jaeckhi Bengtsson, 1989 Scythris lhommei Bengtsson & Passerin d'Entrèves, 1988 Scythris parnassiae Bengtsson, 1997 Scythris popescugorji Passerin d'Entrèves, 1984 Scythris ridiculella Caradja, 1920 Scythris tergestinella (Zeller, 1855) Scythris tumidella K. & T.Nupponen, 2001 alseriella group Scythris alseriella (Turati, 1879) Scythris fasciatella (Ragonot, 1880) Australian group Scythris adelopa Meyrick, 1897 Scythris celidopa Meyrick, 1921 Scythris ceratocosma Meyrick, 1897 Scythris crypsigramma Meyrick, 1897 Scythris detestata Meyrick, 1922 Scythris diatoma (Turner, 1927) Scythris erebospila Meyrick, 1897 Scythris fumida Turner, 1923 Scythris hologramma (Lower, 1899) Scythris leucochyta (Turner, 1947) Scythris paredra Meyrick, 1897 Scythris pleonectis Meyrick, 1897 Scythris plocanota Meyrick, 1897 Scythris praestructa Meyrick, 1922 Scythris rhabducha Meyrick, 1897 Scythris sporadica Meyrick, 1897 Scythris xenonympha (Lower, 1900) bagdadiella group Scythris bagdadiella Amsel, 1949 Scythris caballoides Nupponen, 2009 Scythris kabulella Passerin d'Entrèves & Roggero, 2011 Scythris semifascia Nupponen, 2010 Scythris tugaiensis Nupponen, 2009 bazaensis group Scythris bazaensis Bengtsson, 1997 boseanella group Scythris boseanella Klimesch, 1986 camelella group Scythris camelella Walsingham, 1907 canescens group Scythris angustella Nupponen, 2009 Scythris appendicella Bengtsson, 1997 Scythris articulatella Chrétien, 1915 Scythris barbatella Chrétien, 1915 Scythris biskraensis Bengtsson, 1997 Scythris canescens (Staudinger, 1880) Scythris cupellella Bengtsson, 1997 Scythris curlettii Bengtsson, 1997 Scythris deserticola Nupponen, 2010 Scythris eremella Chrétien, 1915 Scythris eucharis Walsingham, 1907 Scythris falkovitshi Nupponen, 2009 Scythris karvoneni Nupponen, 2010 Scythris maculosa Bengtsson, 1997 Scythris neftae Bengtsson, 1997 Scythris pura Walsingham, 1907 Scythris senecai Bengtsson, 1997 Scythris tessulatella Rebel, 1903 caramani group Scythris albisaxella K. & T.Nupponen, 2000 Scythris arkaimensis Bengtsson, 2000 Scythris caramani Staudinger, 1880 Scythris cervella K. & T.Nupponen, 2001 Scythris cultelloides Nupponen & Sinev, 2011 Scythris eevae Nupponen, 2007 Scythris erinacella Nupponen, 2003 Scythris gorbunovi Nupponen, 2003 Scythris hamatella K. & T.Nupponen, 2001 Scythris hebesella Nupponen, 2005 Scythris kullbergi Bengtsson, 1997 Scythris lagomorphella Nupponen, 2002 Scythris malozemovi Nupponen, 2003 Scythris marashi Passerin d'Entrèves & Roggero, 2012 Scythris sinevi Nupponen, 2003 cicadella group Scythris arenbergeri Passerin d'Entrèves, 1986 Scythris bajanlegi Passerin d'Entrèves & Roggero, 2006 Scythris cicadella (Zeller, 1839) Scythris latilineella K. Nupponen, 2013 Scythris moldavicella Caradja, 1905 (= S. balcanica) Scythris potentillella (Zeller, 1847) Scythris ventosella Chrétien 1907 cistorum group Scythris cistorum (Milliére, 1876) Scythris nieukerkeni Bengtsson, 1989 Scythris rondaensis Bengtsson, 1997 crassiuscula group Scythris crassiuscula (Herrich-Schäffer, 1855) crypta group Scythris crypta Hannemann 1961 decrepidella group Scythris decrepidella Bengtsson, 1997 Scythris goundafae Bengtsson, 1997 dissitella group Scythris dissitella (Zeller, 1847) elenae group Scythris elenae K.Nupponen, 2000 Scythris juerivetei Nupponen, 2005 empetrella group Scythris empetrella Karsholt & Nielsen 1976 Scythris gozmanyi Passerin d'Entrèves 1986 fallacella group Scythris baldensis Passerin d'Entrèves 1979 Scythris fallacella (Schläger 1847) Scythris larzacensis Delmas, 2010 Scythris oelandicella Müller-Rutz 1922 Scythris sappadensis Bengtsson, 1992 Scythris tremalzoi Bengtsson, 1992
formicella group Scythris formicella Chrétien, 1915 Scythris kebiliensis Bengtsson, 1997 Scythris minima Bengtsson, 1997 fuscoaenea group Scythris derrai Bengtsson, 1991 Scythris dissimilella (Herrich-Schäffer, 1855) Scythris fuscoaenea (Haworth, 1828) Scythris parafuscoaenea Bengtsson, 1991 Scythris tabelli Junnilainen, 2002 Scythris tenuivittella (Stainton, 1867) Scythris traugotti Bengtsson, 1991 Scythris vartianae Kasy, 1962 Scythris vittella (O.Costa, 1834) fuscopterella group Scythris fuscopterella Bengtsson 1977 gobiensis group Scythris gobiensis Passerin d'Entrèves & Roggero, 2006 grandipennis group Scythris bornicensis Jäckh, 1977 Scythris constanti Walsingham, 1898 Scythris cupreella (Staudinger, 1859) Scythris ericetella (Heinemann, 1872) Scythris grandipennis (Haworth, 1828) Scythris kasyi Hannemann, 1962 Scythris maroccensis Jäckh, 1977 Scythris nevadensis Passerin d'Entrèves, 1990 Scythris salviella Meess, 1910 Scythris scipionella (Staudinger, 1859) Scythris scorpionella Jäckh, 1977 Scythris siculella Jäckh, 1977 Scythris tabidella (Herrich-Schäffer, 1855) gravatella group Scythris gravatella (Zeller, 1847) iagella group Scythris iagella Chrétien 1925 inclusella group Scythris inclusella (Lederer, 1855) inertella group Scythris ethmiella Amsel, 1974 Scythris inertella (Zeller, 1855) Scythris xanthopygella (Staudinger, 1859) inspersella group Scythris inspersella (Hübner, ) insulella ("ericivorella") group Scythris ericivorella (Ragonot, 1880) Scythris insulella (Staudinger, 1859) karinae group Scythris karinae Bengtsson 1991 karsholti group Scythris karsholti Bengtsson, 1997 klimeschi group Scythris guimarensis Bengtsson, 1997 Scythris klimeschi Passerin d'Entrèves, 1983 knochella group Scythris ambustella Bengtsson, 1997 Scythris aspromontis Jäckh, 1978 Scythris ciliatella Zerny, 1936 Scythris heinemanni Scythris iberica Jäckh, 1978 Scythris knochella (Fabricius, 1794) Scythris reticulella Nupponen, 2010 Scythris schawerdae Rebel, 1931 Scythris setiella (Zeller, 1870) Scythris staudingeri Jäckh, 1978 laminella group Scythris andersi Bengtsson, 1991 Scythris bolognella Jäckh, 1978 Scythris braschiella (O.Hofmann, 1897) Scythris bubaniae Walsingham, 1907 Scythris danilevskyi Nupponen & Sinev, 2011 Scythris eberhardi Bengtsson, 1997 Scythris frankeniella Chrétien, 1916 Scythris kailai Bengtsson, 1997 Scythris laminella (Denis & Schiffermüller, 1775) Scythris saxella Bengtsson, 1991 Scythris thomisioides Bengtsson, 1997 limbella group – possibly subgenus Scythris Scythris elegantella (D.Lucas, 1955) Scythris limbella lobella group Scythris lobella K. Nupponen, 2013 mariannae group Scythris mariannae (Bengtsson, 1997) martini group Scythris bengtbengtssoni Vives, 1994 Scythris martini Bengtsson, 1991 meanderis group Scythris meanderis Bengtsson, 1997 monochreella group Scythris monochreella (Ragonot, 1896) mus group Scythris mus Walsingham, 1898 nigrella group Scythris nigrella Jäckh, 1978 nipholecta group Scythris nipholecta Meyrick, 1924 noricella group Scythris noricella (Zeller, 1843) obscurella group – possibly subgenus Galanthia Scythris amphonycella (Geyer, ) Scythris bengtssoni Patocka & Liska, 1989 Scythris cuspidella (Denis & Schiffermüller, 1775) Scythris flavilaterella (Fuchs, 1886) Scythris hornigii (Zeller, 1855) Scythris lampyrella (Constant, 1865) Scythris mediella (Constant, 1855) Scythris obscurella (Scopoli, 1763) Scythris rouxella (Constant 1865) (sometimes in C. hornigii, tentatively placed here) Scythris speyeri (Heinemann & Wocke, 1876) Scythris shafferi Bengtsson, 2005 Scythris abruptella Sachkov, 2013 palustris group Scythris palustris (Zeller, 1855)
pascuella group Scythris aciella Bengtsson, 1997 Scythris albidella (Stainton, 1867) Scythris albosuffusella Nupponen, 2007 Scythris barguzinensis Bengtsson & Liska, 1996 Scythris bidzilyai Sachkov, 2013 Scythris bifissella (O.Hofmann, 1889) Scythris caroxylella Falkovitsh, 1969 Scythris claudiae Passerin d'Entrèves & Roggero, 2011 Scythris cornuella Bengtsson, 1997 Scythris cramella Nupponen, 2010 Scythris flavidella Preissecker, 1911 Scythris fluxilis Falkovitsh, 1986 Scythris hemicycliella Nupponen, 2005 Scythris kantarae Bengtsson, 1997 Scythris karinupponei Bengtsson, 2000 Scythris lagunae Jäckh, 1978 Scythris mikkolai Sinev, 1993 Scythris nigridorsella Nupponen, 2007 Scythris orientella Sinev, 2001 Scythris pascuella (Zeller, 1855) Scythris paullella (Herrich-Schäffer, 1855) Scythris perlucidella K. & T.Nupponen, 2000 Scythris pudorinella (Möschler, 1866) Scythris rotundella Nupponen, 2010 Scythris satyrella Staudinger, 1880 Scythris subaerariella (Stainton, 1867) Scythris tabescentella (Staudinger, 1880) passerini group Scythris passerini (Bengtsson, 1997) penicillata group Scythris buraetica Nupponen, 2007 Scythris penicillata (Chrétien 1900) Scythris spinella K. & T.Nupponen, 2001 petrella group Scythris arachnodes Walsingham, 1908 Scythris hierroella Klimesch, 1986 Scythris petrella Walsingham, 1908 Scythris pseudoarachnodes Bengtsson, 1997 picaepennis group Scythris disparella (Tengström, 1848) Scythris picaepennis (Haworth, 1828) pinker group Scythris pinkeri Klimesch, 1986 platypyga group Scythris platypyga (Staudinger, 1880) podoliensis group Scythris podoliensis Rebel, 1938 polycarpaeae group Scythris polycarpaeae Klimesch, 1986 productella group Scythris nielseni Passerin d'Entrèves & Roggero, 2003 Scythris productella (Zeller, 1839) pulicella group Scythris antisymmetrica Nupponen, 2009 Scythris apotomella Nupponen, 2007 Scythris azrouensis Bengtsson, 1997 Scythris corleyi Bengtsson, 1997 Scythris langohri Passerin d'Entrèves, 1990 Scythris pilella Bengtsson, 1991 Scythris pulicella (Staudinger, 1859) Scythris skulei Bengtsson, 1997 Scythris veletae Passerin d'Entrèves, 1990 punctivittella group Scythris albostriata Hannemann, 1961 Scythris apicistrigella (Staudinger, 1870) Scythris atlasensis Bengtsson, 1997 Scythris confluens (Staudinger, 1870) Scythris cycladeae Jäckh, 1978 Scythris emichi (Anker, 1870) Scythris landryi Passerin d'Entrèves & Roggero, 2003 Scythris punctivittella (O.Costa, 1836) Scythris trinacriae Passerin d'Entrèves 1984 rubioi group Scythris rubioi Agenjo, 1962 – possibly subgenus Rubioia schleichiella group Scythris glacialis (Frey, 1870)