Plants

Bennettitales Name Novelty Status Authors Age Unit Location Notes ImagesPterophyllum palmulumSp novvalidMiller & HickeyCretaceousLate AlbianWinthrop FormationA williamsoniaceous leaf morphospecies.Williamsonia aculeataSp novvalidMiller & HickeyCretaceousLate AlbianWinthrop FormationA williamsoniaceous strobilus morphospecies.Williamsonia winthropensisSp novvalidMiller & HickeyCretaceousLate AlbianWinthrop FormationA williamsoniaceous strobilus morphospecies.

Cycadales Name Novelty Status Authors Age Unit Location Notes ImagesAndrostrobus scutulatusSp novvalidMiller & HickeyCretaceousLate AlbianWinthrop FormationA male cycad cone of uncertain familial affinities.Laceriagen et sp novvalidMiller & HickeyCretaceousLate AlbianWinthrop FormationA cycad morphogenus of uncertain affiliation.The type species is L. pulcraOrthoneuragen et sp novvalidMiller & HickeyCretaceousLate AlbianWinthrop FormationA cycad morphogenus of uncertain affiliation.The type species is L. longissimaNilssoniocladus corrugatusSp novvalidMiller & HickeyCretaceousLate AlbianWinthrop FormationA nilsoniaceous leaf morphospecies.

Czekanowkiales Name Novelty Status Authors Age Unit Location Notes ImagesCzekanowskia durabilisSp novvalidMiller & HickeyCretaceousLate AlbianWinthrop FormationA czekanowskiaceous leaf morphospecies

Pinales Name Novelty Status Authors Age Unit Location Notes ImagesAthrotaxis parvistrobiliSp novvalidMiller & HickeyCretaceousLate AlbianWinthrop FormationA cupressaceous species.Brachyphyllum cracenenteSp novvalidMiller & HickeyCretaceousLate AlbianWinthrop FormationA pinophyte species of uncertain affiliation.Conagogen et sp novvalidMiller & HickeyCretaceousLate AlbianWinthrop FormationA pinophyte cone morphogenus of uncertain affiliation.The type species is C. tonsifera Genus also includes C. chartifera, C. fornicata, C. spinosa, & C. tonsiferaElatocladus gracilisSp novvalidMiller & HickeyCretaceousLate AlbianWinthrop FormationA pinophyte species of uncertain affiliation.Geinitzia dentifoliaComb novvalid(Fontaine) Miller & HickeyCretaceousPatuxent FormationA pinophyte species of uncertain affiliation. Moved from Sphenolepidium dentifolium (1889)Also includes Sequoia ambigua , Sphenolepidium pachyphyllum (1889) & Sphenolepidium recurvifolium (1889)Geinitzia subtilisSp novvalidMiller & HickeyCretaceousLate AlbianWinthrop FormationA pinophyte species of uncertain affiliation.Pagiophyllum tenuifoliumSp novvalidMiller & HickeyCretaceousLate AlbianWinthrop FormationA pinophyte species of uncertain affiliation.Conagogen et sp novvalidMiller & HickeyCretaceousLate AlbianWinthrop FormationA pinophyte morphogenus of uncertain affiliation.The type species is P. creberumPodozamites decrescensComb & syn novvalid(Fontaine) Miller & HickeyCretaceousAptianPotomac GroupA pinophyte species of uncertain affiliation. Moved from Nageiopsis decrescens (1889)Also includes Nageiopsis heterophylla (1889) & Nageiopsis microphylla (1889)Podozamites varifoliaSp novvalidMiller & HickeyCretaceousLate AlbianWinthrop FormationA pinophyte species of uncertain affiliation.Sphenolepis conditaComb novvalid(Lesquereaux) Miller & HickeyCretaceousDakota FormationA cupressaceous species.Moved from Sequoia condita (1876)

Pteridospermopsida Name Novelty Status Authors Age Unit Location Notes ImagesSagenopteris pandaSp novvalidMiller & HickeyCretaceousLate AlbianWinthrop FormationA caytoniaceous pteridospermopsid leaf morphospecies

Angiosperms

Name Novelty Status Authors Age Unit Location Notes ImagesCornus piggaeSp novValidManchester, Xiang, and XiangTiffanianSentinel Butte FormationOldest member of Cornus subgenus Cornus.Hymenaea allendissp novValidCalvillo-Canadell, Cevallos-Ferriz & Rico-ArceLate Oligocene - Early MioceneMexican amberSecond Hymenaea sp. from Mexican amberLagokarposGen et sp novValidMcMurran & ManchesterLate Paleocene - YpresianGreen River FormationFossil Butte MemberA fruit of uncertain affinity Also found in the Clarno Formation (Oregon) & Horsefly Shales (British Columbia) The type species is L. lacustrisPloufoliaGen. nov.ValidSender et al.AlbianUtrillas FormationSagariaGen et sp novValidBravia, Barone Lumagab, & MickleMiddle AlbianMonti Alburni, near PetinaSolaranthusGen et sp novValidShaolin Zheng & Xin WangMiddle JurassicJiulongshan FormationAn early flowering plant.XingxueanthusGen et sp novValidXin Wang & Shijun WangMiddle JurassicHaifanggou FormationAn early flowering plant.

Other plants Name Novelty Status Authors Age Unit Location Notes ImagesDesmiophyllum truncatumSp novvalidMiller & HickeyCretaceousLate AlbianWinthrop FormationA plant species of uncertain affiliations.

Molluscs

Newly named bivalves Name Novelty Status Authors Age Unit Location Notes ImagesEmiliodontaGen. nov.valid SánchezCaradocDon Braulio FormationReplacement name for Emiliania Sánchez, 1999 preoccupied by Emiliania Hay & Mohler, 1967Erodona doellojuradoiSp. nov.validPérez, Iturerría & GriffinLate MioceneParaná FormationAn erodonid.Polymesoda muravchikiSp. nov.validPérez, Iturerría & GriffinLate MioceneParaná FormationA cyrenid.

Arthropods

Fishes

Amphibians

Newly named amphibians Name Status Authors Age Unit Location Notes ImagesDeltaherpeton Valid Bolt LombardLate Viséan A colosteid.Fedexia Valid Berman Henrici et al.Berman, Henrici, Brezinski, Kollar.GzhelianCasselman Formation A trematopid genus currently among the oldest known vertebrates with a primarily terrestrial lifestyle.Hungarobatrachus Valid Szentesi VenczelSantonianCsehbánya Formation Advanced frog.Madygenerpeton Valid Schoch Voigt BuchwitzMiddle/Upper TriassicMadygen Formation A chroniosuchid reptiliomorph.Pelobates fahlbuschiValid BöhmeMioceneA European spadefoot toad.

Basal reptiles

Newly named basal reptiles

Name Status Authors Age Unit Location Notes ImagesMicroleterValid Tsuji Muller ReiszLower PermianA basal parareptile. The species is M. mckinzieorum. PhonodusValidModesto Scott et al.Modesto, Scott, Botha-Brink, Reisz.Induan (early Triassic)Katberg FormationThe earliest known leptopleuronine procolophonid. ReiszorhinusValidSumida Dodick et al.Sumida, Dodick, Metcalf, Albright.Lower PermianWaggoner Ranch FormationA basal captorhinid. The species is Reiszorhinus olsoni.

Ichthyopterygians

Newly named ichthyopterygians Name Status Authors Age Unit Location Notes ImagesArthropterygius Valid MaxwellQueen Elizabeth Islands A new genus for "Ophthalmosaurus" chirsorum (Russell, 1993).Athabascasaurus Valid Druckenmiller MaxwellLower AlbianClearwater Formation The most complete and stratigraphically oldest known ichthyosaur from the Cretaceous of North America.Barracudasauroides Valid MaischMiddle TriassicGuanling Formation A new genus for "Mixosaurus" panxianensis (Jiang, Schmitz, Hao & Sun, 2006).Mixosaurus xindianensis Valid Chen ChengMiddle TriassicGuanling Formation Omphalosaurus merriami Valid MaischEarly TriassicSticky Keep Formation A species of a possible ichthyopterygian genus Omphalosaurus.

Lepidosauromorphs

Newly named plesiosaurs Name Status Authors Age Unit Location Notes ImagesAlexeyisaurusValid Sennikov ArkhangelskyNorianWilczek FormationA new elasmosauridMeyerasaurusValid Smith VincentLower ToarcianPosidonia ShaleA pliosaur, a new genus for "Plesiosaurus" victor (Fraas, 1910).

Newly named basal lepidosaurs Name Status Authors Age Unit Location Notes ImagesCargniniaValid Bonaparte Schultz et al.Bonaparte, Schultz, Soares, Martinelli.Carnian - NorianCaturrita FormationA basal lepidosaur.

Newly named lizards Name Status Authors Age Unit Location Notes ImagesAdriosaurus skrbinensisValid Caldwell PalciLate CenomanianSkrbinaBavaricordylus molassicusValid BöhmeMiocene (late Karpatian)A cordylid, a species of Bavaricordylus.HeadonhilliaValid Klembara GreenLate Eocene, Ludian (Priabonian)Bembridge Limestone FormationAn anguine lizard. The species is H. parva.Heloderma welcommeiValid Herman Van Den EeckhautEoceneAn anguimorph lizard. Herman and Van Den Eeckhaut (2010) consider it to be a species of Heloderma (though the authors define the genus Heloderma more broadly than most herpetologists, and explicitly synonymize the glyptosaurine genus Placosaurus with it).KleskunsaurusValid Nydam Caldwell FantiUpper CampanianWapiti FormationA scincomorph lizard. The species is K. grandeprairiensis.LiushusaurusValid Evans WangLower CretaceousYixian FormationA scincogekkonomorph lizard. The species is L. acanthocaudata.PedrerasaurusValid Bolet EvansEarly Cretaceous (late Berriasian-early Valanginan)Montsec's Formation (ca)A scincogekkonomorph lizard. The species is P. latifrontalis.Tropidophorus bavaricusValid BöhmeMiocene (late Karpatian)A lygosomine skink, a species of Tropidophorus.Varanus debieiValid Herman Van Den EeckhautEoceneA monitor lizard.

Newly named snakes Name Status Authors Age Unit Location Notes ImagesColombophis spinosusValid Hsiou, Albino & FerigoloLate MioceneSolimões FormationAn alethinophidian snake, a species of Colombophis.KelyophisValid Laduke Krause Scanlon KleyLate Cretaceous (Maastrichtian)Maevarano FormationA nigerophiid snakeMenaranaValid Laduke Krause Scanlon KleyLate Cretaceous (Maastrichtian)Maevarano FormationA madtsoiid snakeMicronatrixValid Parmley & HunterLate Miocene (Clarendonian)A natricine colubrid snake. The type species is Micronatrix juliescottae.Sanajeh Valid Wilson Mohabey et al.Wilson, Mohabey, Peters, Head.MaastrichtianLameta FormationA madtsoiid snake which preyed on hatchling sauropods.

Turtles

Newly named turtles

Name Status Authors Age Unit Location Notes ImagesCerrejonemysValid Cadena Bloch JaramilloPaleoceneCerrejón FormationChupacabrachelysValid Lehman WickCampanianAguja FormationA taphrosphyini bothremydidGamerabaenaValid Lyson JoyceLate Cretaceous (Maastrichtian)Hell Creek FormationItilochelysValid Danilov Averianov YarkovDanianBeryozovaya bedsJiangxichelysValid Tong MoLate CretaceousGanzhouA nanhsiungchelyid turtle.LiaochelysValid ZhouEarly CretaceousJiufotang FormationA macrobaenid turtle.MexichelysValid Parham PyensonLate CampanianCerro del Pueblo FormationA sea turtle. A new genus for "Euclastes" coahuilaensis (Brinkman et al., 2009).PacifichelysValid Parham PyensonMiocene A sea turtle.Pangshura tatrotiaValid Joyce LysonPlioceneTatrot FormationA species of Pangshura."Trionyx" kansaiensisValid Vitek & DanilovLate CretaceousA trionychid with unclear systematic position, a species of Trionyx sensu lato.

Archosauromorphs

Newly named basal archosauromorphs Name Status Authors Age Unit Location Notes ImagesAzendohsaurus madagaskarensisValid Flynn Nesbitt et al.Flynn, Nesbitt, Parrish, Ranivoharimanana, Wyss.Ladinian/CarnianMakay Formation A basal archosauromorph.BentonyxValid Langer Montefeltro et al.Langer, Montefeltro, Hone, Whatley, Schultz.Middle TriassicOtter Sandstone Formation A rhynchosaur.KoilamasuchusValid Ezcurra Lecuona MartinelliEarly TriassicQuebrada de los Fósiles Formation The best-known basal archosauriform from South America.Megalancosaurus endennaeValid Renesto Spielmann et al.Renesto, Spielmann, Lucas, Tarditi Spagnoli.Alaunian (early Norian)Zorzino Limestone Formation A drepanosaurid that is known from MBSN 25, a partial skeleton (Partial postcranial skeleton).TeyumbaitaValid Montefeltro Langer SchultzLate TriassicCaturrita Formation Ischigualasto Formation A rhynchosaur. A new genus for "Scaphonyx" sulcognathus (Azevedo & Schultz, 1987).Uatchitodon schneideriValid Mitchell Heckert SuesLate TriassicCumnock Formation Bluewater Creek Formation A venomous archosauriform known only from isolated teeth.Vallesaurus zorzinensisValid Renesto Spielmann et al.Alaunian (early Norian)Zorzino Limestone Formation A basal drepanosauromorph that is known from MCSNB 4783, a set of vertebrae and hindlimbs.

Archosaurs

Synapsids

Newly named non-mammalian synapsids Name Status Authors Age Unit Location Notes ImagesBeishanodonValid Gao Fox