Orthalicoidea is a superfamily of air-breathing land snails, terrestrial gastropod mollusks in the infraorder Orthalicoidei of the suborder Helicina.MolluscaBase eds. (2021). MolluscaBase. Simpulopsidae Schileyko, 1999. Accessed through: World Register of Marine Species at: http://www.marinespecies.org/aphia.php?p=taxdetails&id=870047 on 2021-02-05
Distribution The Orthalicoidea is a dominant faunal element in the Neotropics, but also has a number of genera with a Gondwanan distribution.
Taxonomy
2005 taxonomy This taxonomy, as accepted by Bouchet & Rocroi, was based on the study by Nordsieck, published in 1986, and the classification as formulated by Schileyko in 1999. However Bouchet & Rocroi diverged from this classification in uniting the families Bulimulidae and Orthalacidae. They also considered the family Placostylidae as a distinct family from Orthalicidae, and the family Coelociontidae a distinct family from Urocoptidae. However, the position of the families Megaspiridae and Grangerellidae is doubtful.
The following seven families have been recognized in the taxonomy of Bouchet & Rocroi (2005):
Family Cerionidae Pilsbry, 1901 Family Coelociontidae Iredale, 1937 † Family Grangerellidae Russell, 1931 Family Megaspiridae Pilsbry, 1904 Family Orthalicidae Albers, 1860 Family Placostylidae Pilsbry, 1946 Family Urocoptidae Pilsbry, 1898 (1868) Vidaliellidae H. Nordsieck, 1986 †
2008 taxonomy Uit de Weerd moved two families Urocoptidae and Cerionidae to newly established superfamily Urocoptoidea based on molecular phylogeny research in 2008.Uit de Weerd D. R. (2008). "Delimitation and phylogenetics of the highly diverse land snail family Urocoptidae (Gastropoda, Pulmonata) based on 28S rRNA sequence data: A reunion with Cerion". Journal of Molluscan Studies 74: 317-329. .
2009 taxonomy Family Prestonellidae van Bruggen, 1978 - A study by Herbert and Mitchell, published in 2009, places this enigmatic family, of which the phylogenetic relationships were previously unknown, in the Gondwanan superfamily Orthalicoidea, while it was previously tentatively placed as a synonym of Aillyidae.
2010 taxonomy Breure et al. (2010) moved Prestonella and Bothriembryon to Placostylidae and they elevated Bulimulinae to Bulimulidae, Odontostomini to Odontostomidae, Amphibuliminae to Amphibulimidae. They also removed Coelociontidae from Orthalicoidea.Breure A. S. H., Groenenberg D. S. J. & Schilthuizen M. (2010). "New insights in the phylogenetic relations within the Orthalicoidea (Gastropoda, Stylommatophora) based on 28S sequence data". Basteria 74(1-3): 25-31.
2012 taxonomy Breure & Romero (2012) confirmed previous results from 2010, additionally they elevated Simpulopsini to Simpulopsidae, renamed Placostylidae to Bothriembryontidae.Breure A. S. H. & Romero P. (2012). "Support and surprises: molecular phylogeny of the land snail superfamily Orthalicoidea using a three-locus gene analysis with a divergence time analysis and ancestral area reconstruction (Gastropoda: Stylommatophora)". Archiv für Molluskenkunde: International Journal of Malacology 141(1): 1-20. . Therefore, there are recognized seven extant families and one extinct family within Orthalicoidea: Amphibulimidae P. Fischer, 1873 Bothriembryontidae Iredale, 1937 Bulimulidae Tryon, 1867 † Grangerellidae Russell, 1931 Megaspiridae Pilsbry, 1904 Odontostomidae Pilsbry & Vanatta, 1898 Orthalicidae Albers, 1860 Simpulopsidae Schileyko, 1999 † Vidaliellidae H. Nordsieck, 1986
References This article incorporates CC-BY-3.0 text from the referenceBreure A. S. H. (2011). "Annotated type catalogue of the Orthalicoidea (Mollusca, Gastropoda) in the Royal Belgian Institute of Sciences, Brussels, with description of two new species". ZooKeys 101: 1-50. .
Further reading Borrero F. J. & Breure A. S. H. (Submitted). "The Orthalicoidea (Mollusca: Gastropoda: Stylommatophora) from Colombia and adjacent areas: a review of taxonomy and biogeography. 1. Dryptus Albers, 1860 and Plekocheilus Guilding, 1828".
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