The Chañares Formation is a Carnian-age geologic formation of the Ischigualasto-Villa Unión Basin, located in La Rioja Province, Argentina. It is characterized by drab-colored fine-grained volcaniclastic claystones, siltstones, and sandstones which were deposited in a fluvial to lacustrine environment. The formation is most prominently exposed within Talampaya National Park, a UNESCO World Heritage Site within La Rioja Province.
The Chañares formation is the lowermost stratigraphic unit of the Agua de la Peña Group, overlying the Tarjados Formation of the Paganzo Group, and underlying the Los Rastros Formation. Though previously considered Ladinian in age, U-Pb dating has determined that most or all of the Chañares Formation dates to the early Carnian stage of the Late Triassic.Kent et al, 2014, p.7959
The Chañares Formation has provided a diverse and well-preserved faunal assemblage which has been studied intensively since the 1960s. The most common reptiles were proterochampsids (Chanaresuchus, Tropidosuchus, and Gualosuchus), which lived alongside true archosaurs such as Lewisuchus, Lagerpeton, Marasuchus, Gracilisuchus, and Luperosuchus. Cynodonts were abundant, represented by the medium-sized traversodontid Massetognathus, as well as smaller carnivores such as Chiniquodon and Probainognathus. The largest animal in the ecosystem was the giant dicynodont Dinodontosaurus. An older faunal assemblage, distinguished by the large erpetosuchid Tarjadia, has been discovered in the earliest part of the formation. The formation as a whole is considered one of the best sources of Carnian-age tetrapods in South America, along with the slightly younger Ischigualasto Formation which lies above the Los Rastros Formation.
Geology The Chañares Formation is the lowermost unit of the Agua de la Peña Group, representing the onset of the first syn-rift phase of the Ischigualasto-Villa Unión Basin. It unconformably overlies red fluvial (river) sediments of the Tarjados Formation of the Paganzo Group, and is conformably overlain by greenish lake and delta sediments of the Los Rastros Formation. The Chañares Formation has also had an interesting history with the lower part of the Ischichuca Formation. This sequence of sediments on the western edge of the basin has occasionally been considered to take priority over the Chañares Formation and the lower part of the Los Rastros Formation. However, this is rare due to the historical significance of the Chañares Formation. As a result, some authors restrict the Ischichuca Formation to a few layers of lake and delta sediments between the Chañares and Los Rastros formations, while others reject using it in the first place.
The Chañares formation was originally thought to be deposited during the Ladinian stage of the Middle Triassic. However, Uranium-Lead radiometric dating by Marsicano et al. (2016) later found that a large portion of the formation was deposited in the early Carnian (237–234 Ma), near the start of the Late Triassic.Marsicano et al., 2016 2020 U-Pb dating of the overlying lower Los Rastros Formation yielded an age of 234.47 ± 0.44 Ma, making the vast majority of the Chañares Formation lowermost Carnian.Mancuso et al., 2020 Nevertheless, the Ladinian-Carnian boundary may still lie within the first few meters of the formation, despite the primary fossiliferous sections being well-supported as early Carnian in age.
The formation is primarily sandstone, siltstone, and claystone, arranged in a specific sequence of facies. A distinct and uneven unconformity separates the base of the Chañares Formation from the underlying Tarjados Formation. In the field, this unconformity can be identified by a high-relief layer of chert at the top of the Tarjados Formation.Rogers et al., 2001Mancuso et al., 2014Ezcurra et al., 2017Pérez et al., 2018
Fluvial olive-grey facies Above this unconformity lies the oldest section of the Chañares Formation, a package of olive-grey fluvial sediments. As one goes up the section, increasingly finer beds of sandstone and siltstone are interlayered with coarser lenses, corresponding to periodic sheet floods along braided rivers. Weakly-developed palaeosols can be found within this section, filled with root traces, pebbles and small brown calcareous nodules.Fiorelli et al., 2018 Winding systems of burrows have also been found in this section, likely created by small cynodonts.
Though fossils are relatively uncommon in the lower fluvial beds of the Chañares Formation, they are taxonomically and taphonomically distinct from those of succeeding layers. The most common remains are from large dicynodonts (possibly referable to Dinodontosaurus), the large erpetosuchid Tarjadia, and small non-massetognathine cynodonts closely related to Aleodon and Scalenodon. This ecosystem has been termed the Tarjadia Assemblage Zone, in order to distinguish it from the slightly younger classic Chañares assemblage. No radiometric dating has been done on this section, but it may contain the Ladinian-Carnian boundary based on dates obtained within the younger facies.
Fossiliferous bluish facies Above the olive-grey fluvial beds is the most fossiliferous and well-studied portion of the formation. This section is characterized by wide and massive layers of very fine bluish-grey sandstone, siltstone, and claystone. These layers have a high concentration of volcanic ash and debris, ranging from glassy shards at the base to weathered bentonites at the top. Nearby volcanic eruptions likely impacted the local climate and river systems, shifting the depositional regime from a stable braided fluvial system to one dominated by shallow floodplains and lahars. Marsicano et al. (2016) obtained a CA-TIMS U-Pb age of 236.1 ± 0.6 Ma from a siltstone bed immediately below the first major fossil layer within the bluish facies. Ezcurra et al. (2017) later studied a slightly older sandstone bed using LA-MC-IMP-MS U-Pb dating. They found a date of 236.2 ± 1.1 Ma from a cluster of the three youngest zircons.
The most productive and historically relevant fossil beds of the Chañares Formation lie within these volcaniclastic layers. The layers are replete with massive calcareous concretions, some up to 2 meters in width. They commonly preserve articulated skeletal material, often complete skeletons from multiple taxa in a single concretion. Well-preserved fossils found within concretions typically represent smaller taxa, which were buried rapidly after death. Volcanic catastrophes such as lahars, ash falls, or pyroclastic flows are the preferred cause of these rapid mass mortality events. More fragmentary fossils are also occasionally found outside of concretions, much like fossils of the underlying Tarjadia AZ. Fossils found outside concretions are typically from large animals, and were probably buried more slowly (albeit still fairly rapidly) by gradual processes.
The classic Chañares fauna, recently codified as the Massetognathus-Chanaresuchus Assemblage Zone, is characteristic of this section of the formation. The largest components of the ecosystem include dicynodonts (namely Dinodontosaurus) and indeterminate carnivorous paracrocodylomorphs, similar to taxa from the underlying Tarjadia AZ. By far the most common fossils belong to three species of cynodonts: Massetognathus pascuali, Probainognathus jenseni, and Chiniquodon theotonicus.These three species comprise nearly 3/4ths of the fossils found in the formation, with almost half of the total fossils in the formation referred to Massetognathus alone. Reptile fossils are less prevalent but more diverse, with the most common belonging to the proterochampsid archosauriform Chanaresuchus. Other notable taxa include Lagerpeton, Lagosuchus, and Lewisuchus, which were among the oldest ornithodirans, elaborating on the ancestry of dinosaurs and pterosaurs. Several communal latrines are known from the bluish facies, preserving dicynodont coprolites filled with plant remains.
Upper member The olive-grey fluvial section and the bluish-grey volcaniclastic section collectively form the lower member of the Chañares Formation. They are overlain by an upper member, which is practically devoid of body fossils. The upper member is generally similar to the bluish facies in appearance, with wide beds of fine-grained volcaniclastic sediments with a pale grey color. Unlike the bluish facies, concretions and body fossils are absent, having been replaced by Taenidium worm burrows. These burrows likely indicate that the environment had transitioned into a lacustrine (lake) ecosystem by the time that the upper member was deposited. Some outcrops preserve a few sandstone and conglomerate beds near the top of the upper member. These indicate that the depositional regime continued to shift towards the deltaic environment of the overlying Los Rastros Formation. The coarser beds also contain a few rare body fossils from fish and tetrapods.Gouiric-cavalli et al., 2017 A white tuff near the base of the upper member has been dated to 233.7 ± 0.4 Ma, while a zircon cluster from a sandy tuff at the top of the formation was dated to 233.6 ± 1.1 Ma.
Paleobiota Synapsids Tetrapod burrows, likely produced by small eucynodonts, have been described from the lower section of the formation. A large dicynodont fossil preserves taphonomic markers which help to reconstruct the sequence of burial, decay, and fossilization responsible for articulated fossils in the formation. Cynodonts Cynodonts of the Chañares Formation Genus / Taxon Species Assemblage Zone Material Notes Images Aleodontinae indet. Tarjadia AZ A single partial skull A small indeterminate aleodontine chiniquodontid cynodont, formerly referred to cf. Aleodon. Chiniquodon C. theotonicus Massetognathus-Chanaresuchus AZ Numerous specimens A common chiniquodontid cynodontRiojanodonR. nenoiTarjadia AZTwo partial lower jawsAn aleodontine chiniquodontid cynodont.Massetognathus M. major Massetognathus-Chanaresuchus AZ A single skull A junior synonym of Massetognathus pascualiAbdala & Giannini, 2000 M. pascualiRomer, 1967, III Massetognathus-Chanaresuchus AZ Numerous specimens (almost half of all recovered fossils) An abundant massetognathine traversodontid cynodont M. teruggii Massetognathus-Chanaresuchus AZ Several skulls A junior synonym of Massetognathus pascuali Megagomphodon M. oligodens Massetognathus-Chanaresuchus AZ A pair of skulls A junior synonym of Massetognathus pascuali Pontognathus P. ignotus Massetognathus-Chanaresuchus AZ Partial skull A rare traversodontid cynodont ProbainognathusRomer, 1971, VI P. jenseni Massetognathus-Chanaresuchus AZ Numerous specimens A very common probainognathid cynodontProbelesodon P. lewisiRomer, 1971, V Massetognathus-Chanaresuchus AZ Numerous specimens A probable junior synonym of Chiniquodon theotonicusAbdala & Giannini, 2002 P. minorRomer, 1971, XVIII Massetognathus-Chanaresuchus AZ Numerous specimens A probable junior synonym of Probelesodon lewisi or Chiniquodon theotonicus cf. Scalenodon/Mandagomphodon Tarjadia AZ Two partial jaws A small basal traversodontid cynodont
Dicynodonts Dicynodonts of the Chañares Formation Genus / Taxon Species Assemblage Zone Material Notes Images Dinodontosaurus D. brevirostris Massetognathus-Chanaresuchus AZ Several specimensde los Angeles Ordonez et al., 2020 A large kannemeyeriiform dicynodont D. platyceps Massetognathus-Chanaresuchus AZ Two specimens A massive kannemeyeriiform dicynodont D. platygnathus Massetognathus-Chanaresuchus AZ One fragmentary specimen A massive kannemeyeriiform dicynodont D. sp.Massetognathus-Chanaresuchus AZ Several specimens A massive kannemeyeriiform dicynodont Kannemeyeriiformes indet. Tarjadia AZ Several specimens Indeterminate remains from massive kannemeyeriiform dicynodonts, possibly referrable to Dinodontosaurus StahleckeriaMancuso & Irmis, 2019 S. sp. Tarjadia AZ An incomplete ulna A large stahleckeriine dicynodont
Reptiles