The Cerithioidea is a superfamily of marine, brackish water and freshwater gastropod containing more than 200 genera. The Cerithioidea are included unassigned in the subclass Caenogastropoda. The original name of this superfamily was Cerithiacea, in keeping with common superfamily endings at the time.
Morphology Many cerithioids, as well as some former cerithioids now assigned to other superfamilies, have a "cerithiform" shell. This is a high-spired and regularly coiled shell with a small aperture and a well-developed anterior canal/sinus/notch. There are often strong spiral rows of knobs and tubercles, or axial ridges.
Some families have different shell morphologies. Planaxidae have a lower spire than typical cerithioids. Siliquariidae are uncoiled/irregularly coiled. Turritellidae lack the well-developed anterior canal.
Internally, cerithioids share a number of features, though some are also present in other superfamilies. These are a complex midgut, open pallial gonoducts, aphallate males (i.e. lacking the male copulatory organ), glandular ovipositors and epiathroid/dialyneurous nervous systems.
Ecology Cerithioidea is a very diverse superfamily. Its species can be found worldwide mainly in tropic and subtropic seas on rocky intertidal shores, seagrass beds and algal fronds, but also in estuarine and freshwater habitats. The freshwater species are found on all continents, except Antarctica. They are dominant members of mangrove forests, estuarine mudflats, fast-flowing rivers and placid lakes.Healy J. M. & Wells F. E. (). Mollusca, The Southern Syntthesis. Fauna of Australia. Melbourne, CSIRO publishing. 707 pp.
Fossil record Their fossil record of this superfamily can be traced back as far as the early TriassicTracey S., Todd J. A. & Erwin D. H. (1993). The Fossil Record. London, Chapman & Hall. pages 131–167. but they began radiating mainly during the Cretaceous.Houbrick R. S. (1988). "Prosobranch Phylogeny". Malacological Review, Supplement 4: 88–128.
Taxonomy The Cerithioidea are presumed to be monophyletic (one lineage). However the phylogenetic relationships between its families are still under investigation because mitochondrial recombinant DNA sequences failed to resolve these questions.
2005 taxonomy According to the Taxonomy of the Gastropoda (Bouchet & Rocroi, 2005), the following families are included in Cerithioidea: Amphimelaniidae P. Fischer & Crosse, 1891 Batillariidae Thiele, 1929 - monophyletic † Brachytrematidae Cossmann, 1906 † Cassiopidae Beurlen, 1967 Cerithiidae Fleming, 1822 - monophyletic Dialidae Kay, 1979 Diastomatidae Cossman, 1894 † Eustomatidae Cossmann, 1906 † Ladinulidae Bandel, 1992 † Lanascalidae Bandel, 1992 Litiopidae Gray, 1847 † Maoraxidae Bandel, Gründel, Maxwell, 2000 Melanopsidae H. Adams & A. Adams, 1854 - freshwater snails, polyphyletic Modulidae P. Fischer, 1884 Pachychilidae P. Fischer & Crosse, 1892 Paludomidae Stoliczka, 1868 Pelycidiidae Ponder & Hall, 1983 Pickworthiidae Iredale, 1917 Planaxidae Gray, 1850 Pleuroceridae P. Fischer, 1885 (1863) - freshwater snails, polyphyletic † Popenellidae Bandel, 1992 Potamididae H. Adams & A. Adams, 1854 - monophyletic † Procerithiidae Cossmann, 1906 - If the genus Argyropeza is placed in the Procerithiidae, then this family is no longer exclusively a family of fossils. † Propupaspiridae Nützel, Pan & Erwin, 2002 † Prostyliferidae Bandel, 1992 Scaliolidae Jousseaume, 1912 - monophyletic Semisulcospiridae J. P. E. Morrison, 1952 Siliquariidae Anton, 1838 Thiaridae Gill, 1871 (1823) - freshwater snails, polyphyletic Turritellidae Lovén, 1847 - monophyletic Zemelanopsidae Neiber & Glaubrecht, 2019
(Extinct taxa indicated by a dagger, †.)
It is possible that a further detailed examination may show that the polyphyletic families Melanopsidae and Pleuroceridae are one family. There is also a close phylogenetic relationship between the families Modulidae and Potamididae and between the families Cerithiidae and Litiopidae.
2006 taxonomy Bandel (2006)Bandel K. (2006). "Families of the Cerithioidea and related superfamilies (Palaeo-Caenogastropoda; Mollusca) from the Triassic to the Recent characterized by protoconch morphology - including the description of new taxa". Freiberger Forschungshefte C 511: 59–138. PDF. made numerous changes in Cerithioidea. He classified superfamily Cerithioidea in the clade Cerithimorpha.
Changes include:
superfamily Cerithioidea family Bittiidae Cossmann, 1906 - consider Bittiidae in its own family level. It was as subfamily Bittiinae within Cerithiidae by Bouchet & Rocroi, 2005. family † Maturifusidae - moved to Cerithioidea from Hypsogastropoda family † Canterburyellidae - moved to Cerithioidea from unallocated Sorbeoconcha family † Prisciphoridae - moved to Cerithioidea from unallocated Sorbeoconcha family † Zardinellopsidae Bandel, 2006 - new family family Pachymelaniidae - considered as valid family. It was as synonym of Thiaridae. some Pyrguliferidae members (a synonym) are in Paludomidae and some are in Paramelaniidae (instead of Thiaridae) family Paramelaniidae at family level (instead of a synonym of Paludomidae) and some moves to other taxa
2009 taxonomy Subfamily Semisulcospirinae within Pleuroceridae was elevated to family level under the name Semisulcospiridae by Strong & Köhler (2009).
2017 Taxonomy In the updated taxonomy by Bouchet et al. (2017)are listed below: Batillariidae † Brachytrematidae Cerithiidae Dialidae Diastomatidae † Eustomatidae Hemisinidae P. Fischer & Crosse, 1891 † Ladinulidae Litiopidae † Maoraxidae Melanopsidae † Metacerithiidae Modulidae Pachychilidae Paludomidae Pelycidiidae Pickworthiidae Planaxidae Pleuroceridae † Popenellidae Potamididae † Procerithiidae † Propupaspiridae † Prostyliferidae Scaliolidae Semisulcospiridae Siliquariidae Thiaridae Turritellidae
Unassigned: Microstilifer Warén, 1980
The following two extinct families were moved out: Lanascalidae † Metacerithiidae †
References
External links Halder K. & Sinha P. (2014). "Some Eocene Cerithioids (Gastropoda, Mollusca) from Kutch, Western India, and Their Bearing on Palaeobiogeography of the Indian Subcontinent". Paleontology Journal 2014: Article ID 673469, 11 pp. .