Cochliopidae is a family of small freshwater snails with gills and an operculum, aquatic gastropod mollusks.
Paludestrina d'Orbigny, 1840 is an archaic synonym, and has been placed on the Official Index of Rejected and Invalid Names by ICZN Opinion 2202.MolluscaBase eds. (2021). MolluscaBase. Cochliopidae Tryon, 1866. Accessed through: World Register of Marine Species at: http://www.marinespecies.org/aphia.php?p=taxdetails&id=411706 on 2021-05-13
This family is in the superfamily Truncatelloidea and in the order Littorinimorpha (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005).
Description Cochliopidae snails are characterized by sharp, elongated spire-like structures, and can be found in rivers or brackish water.
2005 taxonomy The family Cochliopidae consists of 3 subfamilies (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005): Cochliopinae Tryon, 1866 - synonyms: Mexithaumatinae D. W. Taylor, 1966, Paludiscalinae D. W. Taylor, 1966 Littoridininae Thiele, 1928 Semisalsinae Guiusti & Pezzoli, 1980 - synonym: Heleobiini Bernasconi, 1991
Genera Liu et al. (2001) have recognized 34 genera with more than 260 species within the subfamily Cochliopinae.
Strong et al. (2008)Strong E. E., Gargominy O., Ponder W. F. & Bouchet P. (2008). "Global Diversity of Gastropods (Gastropoda; Mollusca) in Freshwater". Hydrobiologia 595: 149-166. . have recognized 246 freshwater species within Cochliopidae.
Genera within the family Cochliopidae include:
subfamily Cochliopinae Aphaostracon F. G. Thompson, 1968 Coahuilix Taylor, 1966 Cochliopa Stimpson, 1865 - type genus of the family Cochliopidae Cochliopina Morrison, 1946 Emmericiella Pilsbry, 1909 Lobogenes Pilsbry & Bequaert, 1927 Mexithauma D. W. Taylor, 1966 Nanivitrea Thiele, 1927 Subcochliopa J. P. E. Morrison, 1946 Tepalcatia F. G. Thompson & Hershler, 2002
subfamily Littoridininae Antrobia Hubricht, 1971 - with the only species Antrobia culveri Hubricht, 1971Kabat A. R. & Hershler R. (1993). "The prosobranch snail family Hydrobiidae (Gastropoda: Rissooidea): review of classification and supraspecific taxa". Smithsonian Contributions to Zoology 547: 1-94. PDF. - Tumbling Creek cavesnail Littoridina Souleyet, 1852 Pseudocaspia Starobogatov, 1972 Pseudotryonia Hershler, 2001 Pseudotryonia adamantina Pseudotryonia alamosae Pseudotryonia brevissima Pseudotryonia grahamae Pseudotryonia mica Hershler, Liu & Landye, 2011Hershler R., Liu H.-P. & Landye J. J. (2011). "Two new genera and four new species of freshwater cochliopid gastropods (Rissooidea) from northeastern Mexico". Journal of Molluscan Studies 77(1): 8-23. . Pseudotryonia pasajae Hershler, Liu & Landye, 2011
subfamily Semisalsinae Heleobia Stimpson, 1865Kroll O., Hershler R., Albrecht C., Terrazas E. M., Apaza R., Fuentealba C., Wolff C. & Wilke T. (2012). "The endemic gastropod fauna of Lake Titicaca: correlation between molecular evolution and hydrographic history". Ecology and Evolution 2(7): 1517-1530. . Heleobops Thompson, 1968 Paludiscala D. W. Taylor, 1966 Semisalsa Radoman, 1974
subfamily ? (either Cochliopinae or Littoridininae) Aroapyrgus H. B. Baker, 1931 Balconorbis Hershler & Longley, 1986 † Carinulorbis Yen, 1949 Chorrobius Hershler, Liu & Landye, 2011 Dyris Conrad, 1871 - it has extant species and also 26 species in Miocene Pebas Formation Eremopyprgus Hershler, 1999Hershler R., Liu H.-P. & Landye J. J. (2002). "A new species of Eremopyprgus (Hydrobiidae: Cochliopinae) from the Chihuahuan desert, Mexico: Phylogentic relationships and biogeography". Journal of Molluscan Studies 68: 7-13. PDF.Hershler R. (1999). "A systematic review of the hydrobiid snails (Gastropoda: Rissoidea) of the Great Basin, western United States. Part II. Genera Colligyrus, Fluminicola, Pristinicola, and Tryonia". The Veliger 42(4): 306-337. PDF . † Feliconcha Wesselingh, Anderson & Kadolsky, 2006 - two species from Miocene of the Pebas FormationWesselingh F. P., Anderson L. C. & Kadolsky D. (2006). "Molluscs from the Miocene Pebas Formation of Peruvian and Colombian Amazonia". Scripta Geologica 1333: 19-290. PDF. Feliconcha feliconcha Wesselingh, Anderson & Kadolsky, 2006 Feliconcha reticulata Wesselingh, Anderson & Kadolsky, 2006 (probably extant) Glabertryonia Wesselingh, Anderson & Kadolsky, 2006 - three species † Glabertryonia glabra Wesselingh, Anderson & Kadolsky, 2006 - from Miocene of the Pebas Formation † Glabertryonia sp. 1 - from Pliocene of the Las Piedras Formation (probably extant) Glabertryonia sp. 2 - from Holocene of Surinam, probably extant Juturnia Hershler, Liu & Stockwell, 2002 Lithococcus Pilsbry, 1911 Mesobia F. G. Thompson & Hershler, 1991 Mexipyrgus Taylor, 1966 Minckleyella Hershler, Liu & Landye, 2011 - with the only species Minckleyella balnearis Hershler, Liu & Landye, 2011 Onobops Thompson, 1968 Pyrgophorus Ancey, 1888 Sioliella Haas, 1949 Spurwinkia Davis, Mazurkiewicz & Mandracchia, 1982 Texadina Abbott & Ladd, 1951 Thalassobia Bourguignat in Mabille, 1877
Cladogram A cladogram based on sequences of mitochondrial cytochrome-c oxidase I (COI) genes showing phylogenic relations of species within Cochliopidae:Liu H.-P., Hershler R. & Thompson F. G. (2001). "Phylogenetic Relationships of the Cochliopinae (Rissooidea: Hydrobiidae): An Enigmatic Group of Aquatic Gastropods". Molecular Phylogenetics and Evolution 21(1): 17-25. .
References
Further reading Hershler R., Davis C. L., Kitting C. L & Liu H.-P. (2007). "Discovery of introduced and cryptogenic cochliopid gastropods in the San Francisco Estuary, California". Journal of Molluscan Studies 73(4): 323-332. PDF. Hershler R. & Thompson F. G. (1992). "A Review of the Aquatic Gastropod Subfamily Cochliopinae (Prosobranchia: Hydrobiidae)". Malacological Review, Supplement 5: 1-140. pages 16–18. Wilke T., Haase M., Hershler R., Liu H.-P., Misof B. & Ponder W. (2013) Pushing short DNA fragments to the limit: Phylogenetic relationships of ‘hydrobioid’ gastropods (Caenogastropoda: Rissooidea). Molecular Phylogenetics and Evolution, 66: 715-736
External links
Cochliopidae on WoRMS