Dyakiidae is a family of air-breathing land snails terrestrial pulmonate gastropod mollusks in the superfamily Trochomorphoidea (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005).
Some of the species in this family are sinistral (left-handed) in their shell coiling.
Distribution The family Dyakiidae is endemic to Sundaland in Southeast Asia..
Anatomy In this family, the number of haploid chromosomes is between 26 and 30 (according to the values in this table).Barker G. M. (2001) Gastropods on Land: Phylogeny, Diversity and Adaptive Morphology. 1-146. In: Barker G. M. (ed.) (2001) The biology of terrestrial molluscs. CABI Publishing, Oxon, UK, cited pages: 139-144. .
The family also includes the only known terrestrial gastropods to emit light..
The Digestive system characteristics are as follows. The buccal mass is small. The jaw is smooth. The stomach is very simple with weak muscles (as is the case in the majority of land snails). (These anatomical characteristics also include the family Staffordiidae which was considered part of the Dyakiidae at the time the study was done).
Genera The family Dyakiidae includes the following genera, with a steadily increasing number of described species (71 species until 2007 + 11 new species in 2009.):
Asperitas Gray, 1857 - 14 species Bertia Ancey, 1887 - at least 4 species Dyakia Godwin-Austen, 1891 - type genus, 22 species Elaphroconcha Gude, 1911 - 10 species Everettia Godwin-Austen, 1891Godwin-Austen H. H. (1891) "On a collection of Land-shells made In Borneo by Mr. A. Everett, with Descriptions of supposed new species. Part II. Zonitidae and Helicidae". Proceedings of the Zoological Society of London for the year 1891: 22-47, p. 33. - 25 species Kalamantania Laidlaw, 1931 - only one species: Kalamantania whiteheadi (Godwin-Austen, 1891)Godwin-Austen H. H. (1891) "On a collection of Land-shells made In Borneo by Mr. A. Everett, with Descriptions of supposed new species. Part II. Zonitidae and Helicidae". Proceedings of the Zoological Society of London for the year 1891: 22-47, p. 24. Phuphania Tumpeesuwan, Naggs & Panha, 2007 - 3 species including: Phuphania globosa Tumpeesuwan, Naggs & Panha, 2007Tumpeesuwan C., Naggs F. & Panha S. (31 August 2007) "A new genus and new species of dyakiid snail (Pulmonata: Dyakiidae) from the Phu Phan range, northeastern Thailand". Raffles Bulletin of Zoology 55(2) : 363-369. PDF Pseudoplecta Laidlaw, 1932 - only one species: Pseudoplecta bijuga (Stoliczka, 1873) Pseudoquantula Jirapatrasilp & Panha gen. nov. - only one species: Pseudoquantula lenticularis Jirapatrasilp & Panha sp. nov. Quantula Baker, 1941 - 7 species including: Quantula striata (Gray, 1834) Rhinocochlis Thiele, 1931 - only one species: Rhinocochlis nasuta (Metcalfe, 1852) Sasakina Rensch, 1930 - 5 species
Cladogram The following cladogram shows the phylogenic relationships of this family and superfamily with the other families within the limacoid clade:.
References
Further reading Laidlaw F. F. (1931). "On a new sub-family Dyakiinae of the Zonitidae". Proceedings of the malacological Society of London 19: 190-201. . Schileyko A. A. (2003). "Treatise on recent terrestrial pulmonate mollusks. 10. Ariophantidae, Ostracolethaidae, Ryssotidae, Milacidae, Dyakiidae, Staffordiidae, Gastrodontidae, Zonitidae, Daudebardiidae, Parmacellidae". Ruthenica, Supplement 2. 1309-1466. Bouchet P., Rocroi J.P., Hausdorf B., Kaim A., Kano Y., Nützel A., Parkhaev P., Schrödl M. & Strong E.E. (2017). Revised classification, nomenclator and typification of gastropod and monoplacophoran families. Malacologia. 61(1-2): 1-526
External links Gude, G. K. & Woodward, B. B. (1921). On Helicella, Férussac. Proceedings of the Malacological Society of London. 14 (5/6): 174-190. Londo