Flora Ferns and fern allies NameNoveltyStatusAuthorsAgeType localityLocationNotesImagesMakotopterisGen et sp novvalidStockey, Nishida, & RothwellEoceneYpresianEocene Okanagan HighlandsAllenby FormationPrinceton ChertAn athyriaceous fern The type species is M. princetonensis
Cycads NameNoveltyStatusAuthorsAgeType localityLocationNotesImagesEostangeria pseudopterisSp novvalidKvaček & ManchesterLate Paleoceneupper Fort Union Formation or lower Wasatch FormationAn eostangerioid zamiaceous cycad.Also found in the Eocene John Day Formation of Oregon.
Angiosperms NameNoveltyStatusAuthorsAgeType localityLocationNotesImagesFlorissantia sikhote-alinensiscomb novvalid(Kryshtofovich) ManchesterMioceneAmgu floraA mallow relative, moved from Porana sikhote-alinensis (1921) Quercus hiholensisSp novValidBorgardt & PiggMiddle Miocene"Yakima Canyon Flora"A white oak, permineralized acornsSargentodoxa globosaComb novvalid(Manchester) ManchesterMiddle EoceneClarno FormationA lardizabalaceous species; New comb for Bumelia? globosa (1994)
Arthropods
Insects Name Novelty Status Authors Age Unit Location Notes ImagesCephalotes alveolatusComb novValid(Vierbergen & Scheven)BurdigalianDominican amberA myrmicinae ant, Moved from Zacryptocerus alveolatus (1995)Cephalotes caribicusSp novValidDe Andrade & Baroni UrbaniBurdigalianDominican amberA myrmicinae ant.Cephalotes dieteriSp novValidDe Andrade & Baroni UrbaniBurdigalianDominican amberA myrmicinae ant.Cephalotes hispaniolicusSp novValidDe Andrade & Baroni UrbaniBurdigalianDominican amberA myrmicinae ant.Cephalotes serratusComb novvalid(Vierbergen & Scheven, 1995)BurdigalianDominican amberA myrmicinae ant, moved from Exocryptocerus serratus (1995)Camponotus crozeiSp novValidRiouTurolianMontagne d'AndanceA formicine ant.Electrostephanus neovenatusSp novvalidAguiar & JanzenLutetianBaltic AmberA stephanid waspElectrostephanus sulcatusSp novjr synonymAguiar & JanzenLutetianBaltic AmberA stephanid wasp, Moved to Denaeostephanus sulcatusNeoephemera antiquasp. novvalidSinitchenkovaYpresian Klondike Mountain FormationA neoephemerid mayflyPachycondyla rebekkaesp. novvalidRust & AndersenYpresian Fur FormationA Myrmeciinae ant moved to Ypresiomyrma rebekkae (2006)Penthetria intermediaComb novvalid(Scudder)PriabonianFlorissant Formation A bibionid fly Originally Mycetophaetus intermedius (1892) Moved from Plecia intermedia (1959)
Mollusca
Newly named bivalves Name Novelty Status Authors Age Unit Location Notes ImagesConcavodonta ovalissp novValid SánchezCaradocDon Braulio Formationthird species in the genusConcavodontinaesubfam novValid SánchezOrdovicianone of two Subfamilies in Praenuculidae CuyopsisGen et sp novValid SánchezCaradocDon Braulio FormationMonotypic with the species C. symmetricusEmilianiaGen et sp novhomonym SánchezCaradocDon Braulio Formationgenus name preoccupied by Emiliania Hay & Mohler, 1967, renamed to EmiliodontaHemiconcavodontaGen et sp novValid SánchezCaradocDon Braulio Formationthird genus in the subfamily ConcavodontinaePraenuculinaesubfam novValid SánchezOrdovicianone of two Subfamilies in Praenuculidae Similodonta ceryxSp novValid CopeMiddle OrdovicianGilfach Farm No. 1 Boreholeoldest species in the genus SimilodontaTrigonoconchaGen et sp novValid SánchezCaradocDon Braulio FormationMonotypic with the species T. acutaVillicumiaGen et sp novValid SánchezCaradocDon Braulio FormationMonotypic with the species V. canteraensis
Fish
Newly named actinopterygii ("ray-finned fish") Name Novelty Status Authors Age Unit Location Notes ImagesNardovelifergen et sp novValidC. Sorbini & L. SorbiniCampanianNardo
Reptiles Archosauromorphs Ornithomimid gastroliths documented.Kobayashi et al. (1999). Sanders, Manley, and Carpenter (2001), "Table 12.1" page 167.
Newly named dinosauriforms Name Novelty Status Authors Age Unit Location Notes ImagesEucoelophysisgen et sp novValid Sullivan & LucasNorianChinle Formation, New MexicoSister genus to the Dinosaurs; first identified as a relative of coelophysid.
Newly named dinosaurs Data courtesy of George Olshevsky's dinosaur genera list.
Name Novelty Status Authors Age Unit Location Notes ImagesAchillobatorPerle, A., M. Norell and J. Clark. (1999). A new maniraptoran theropod Achillobator giganticus (Dromaeosauridae) from the Upper Cretaceous of Burkhant, Mongolia. Contributions of the Department of Geology, National University Mongolia (35 pages).Gen et sp novValidPerle, Norell, & ClarkLate CretaceousBayan Shireh FormationA large dromaeosaurid.AgustiniaBonaparte, J.F. (1999). An armored sauropod from the Aptian of Northern Patagonia, Argentina. Proceedings of the Second Gondwanan Dinosaur Symposium (15 October). National Science Museum Monographs 15 (Y. Tomida, T.H. Rich and P. Vickers-Rich, eds.), Tokyo.Gen et sp novValidBonaparteAlbianLohan Cura FormationAn armored sauropod"Airakoraptor"gen novNomen nudum Perle, Norell, & ClarkA dromaeosaurid, now Kuru kulla, which was later described in 2021.AnimantarxCarpenter, K., K. Kirkland, D. Burge, and J. Bird. (1999). "Ankylosaurs (Dinosauria; Ornithischia) of the Cedar Mountain Formation, Utah, and the stratigraphic distributions". In: Vertebrate Paleontology in Utah (D. Gillete, ed.). Utah Geological Survey Miscellaneous Publication 99-1.Gen et sp novValidCarpenter, Kirkland, Burge, & BirdLate CretaceousCedar Mountain Formation, UtahNodosaur whose name means "living citadel.""Archaeoraptor"Czerkas vide Sloan, C.P. (1999). "Feathers for T. rex?" National Geographic 196 (5 - November): pp. 98-107. vide Olson, 2000. Hoax Stephen Czerkas vide:C. P. Sloan vide:OlsonChimera of Yanornis and Microraptor.Atlasaurusgen et sp novValid Monbaron, Russell, & TaquetMiddle JurassicTiougguit FormationA primitive sauropod Beipiaosaurusgen et sp novValidXu, Tang, & WangEarly CretaceousYixian FormationA therizinosaur"Bilbeyhallorum"Burge, D.L., J.H. Bird, B.K. McClelland, and M.A. Cicconetti. (1999). "Comparison of four armored dinosaurs from the Cedar Mountain Formation of Eastern Utah". Journal of Vertebrate Paleontology, Abstracts of Papers, 59th Annual Meeting, Society of Vertebrate Paleontology, Denver, CO, Volume 19, Supplement to n. 3, p. 34A.Nomen nudum Burge, Bird, McClelland, & CicconettiAptianCedar Mountain Formation, Utahinformal name for Cedarpelta. Cedarosaurusgen et sp novValid Tidwell, Carpenter, & BrooksBarremianCedar Mountain Formation, UtahA sauropod.Chaoyangsaurus gen et sp novValid Zhao, Cheng, & XuLate JurassicTuchengzi FormationAn extremely primitive ceratopsianDinheirosaurusgen et sp nov Valid Bonaparte & MateusKimmeridgianAlcobaça FormationA diplodocid."Eugongbusaurus"Knoll, F. (1999). "The family Fabrosauridae". In: IV European Workshop on Vertebrate Palaeontology: p. 65;gen nov Nomen nudum KnollLate JurassicShishugou FormationA ?hypsilophodont that hasn't yet been formally named.GondwanatitanKellner, A.W.A. and S.A.K. de Azevedo. (1999). A new sauropod dinosaur (Titanosauria) from the Late Cretaceous of Brazil. In: Proceedings of the Second Gondwanan Dinosaur Symposium (Y. Tomida, T.H. Rich, and P. Vickers-Rich, eds.): pp. 111-142.gen et sp novValid Kellner & AzevedoLate CretaceousAdamantina FormationA titanosaurGuaibasaurusBonaparte, J.F., J. Ferigolo, and A.M. Ribeiro. (1999). A new early Late Triassic saurischian dinosaur from Rio Grande do Sul State, Brazil. In: Proceedings of the Second Gondwanan Dinosaur Symposium (Y. Tomida, T.H. Rich, and P. Vickers-Rich, eds.): pp. 89-109.gen et sp novValidBonaparte, Ferigolo, & RibeiroLate TriassicCaturrita FormationA sauropodomorph. Jobariagen et sp novValidSereno, et alMiddle JurassicTiourarén FormationA eusauropodLessemsaurusgen et sp novValid BonaparteLate TriassicLos Colorados FormationA primitive sauropodomorphLirainosaurusSanz, J.L., J.E. Powell, J. Le Loueff, R. Martinez, and X. Pereda-Suberbiola. (1999). Sauropod remains from the Upper Cretaceous of Laño (Northcentral Spain). Titanosaur phylogenetic relationships. Est. Mus. Cienc. Nat. de Alava 14 (Num. Espec. 1): pp. 235-255 .gen et sp novValidSanz et alLate Cretaceous Vitoria FormationA titanosaur.Lurdusaurusgen et sp novValid Taquet & RussellAptianElrhaz FormationA hadrosaur "Newtonsaurus"Pickering, S. (1995). A fractal scaling in dinosaurology project (2nd revised printing). Capitola, California: 478 pages; [nomen nudum].gen et sp novNomen nudumWelles vide: Welles & PickeringLate TriassicRhaetic bedsInformal name. Formally describe in 2025. Nigersaurusgen et sp novValid Sereno et alEarly CretaceousElrhaz FormationA rebbachisaurid Nodocephalosaurusgen et sp nov Valid taxonSullivanCampanianKirtland FormationAn ankylosauridProyandusaurusgen novsynonym KnollBajocianShaximiao Formationinformal name for Hexinlusaurus QantassaurusRich, T. H. and P. Vickers-Rich. (1999). The Hypsilophodontidae from southeastern Australia. In: Proceedings of the Second Gondwanan Dinosaur Symposium (Y. Tomida, T.H. Rich, and P. Vickers-Rich, eds.): pp. 167-180.gen et sp novValidRich & Vickers-RichEarly CretaceousWonthaggi FormationA possible hypsilophodontidSantanaraptorKellner, A.W.A. (1999). "Short note on a new dinosaur (Theropoda, Coelurosauria) from the Santana Formation (Romualdo Member, Albian), Northeastern Brazil". Boletim do Museu Nacional (Rio de Janeiro, Brasil), N.S. 9: 8 pages.gen et sp novValid KellnerCenomanianSantana FormationA theropodSaturnaliaLanger, M.C., F. Abdala, M. Richter, and M.J. Benton. (1999). A sauropodomorph dinosaur from the Upper Triassic (Carnian) of southern Brazil. Compte rendu hebdomadaire des séances de l'Académie des Sciences Paris, Sciences de la Terre et des Planètes 329 : pp. 511-517.gen et sp novValid Langer, Abdala, Richter, & BentonCarnianSanta Maria FormationA basal sauropodomorph Sinornithosaurusgen et sp novValid Xu, Wang, & WuAptianYixian FormationA DromaeosauridTangvayosaurusGen et sp novValidAllain et atEarly CretaceousGrès Supérior FormationA titanosaur TehuelchesaurusRich, T.H., P. Vickers-Rich, O. Gimenez, R. Cúneo, P. Puerta, and P. Vacca. (1999). "A new sauropod dinosaur from Chubut Province, Argentina". In: Proceedings of the Second Gondwanan Dinosaur Symposium (Y. Tomida, T.H. Rich, and P. Vickers-Rich, eds.): pp. 61-84.Gen et sp nov ValidRich et alMiddle JurassicCañadon Asfalto FormationA eusauropod
Newly named birds Name Novelty Status Authors Age Unit Location Notes ImagesAmplibuteo concordatus Sp. nov. ValidSteven D. Emslie
Nicholas J. CzaplewskiLate PlioceneFlorida and Arizona:
;An Accipitridae.Anatalavis oxfordi Sp. nov. ValidStorrs L. OlsonEarly EoceneYpresian,
London Clay,
MP 8:An Anseranatidae, Anatalavinae Olson, 1999. In 2002 Jíří Mlíkovský made this species the type species of his not widely accepted new genus Nettapterornis.Aquila bivia Sp. nov. ValidSteven D. Emslie
Nicholas J. CzaplewskiLate PlioceneFlorida:
;An Accipitridae.Argornis caucasicus Gen. nov. et Sp. nov. ValidAleksandr A. KarkhuLate EoceneNorthern CaucasusAn Apodiformes, Jungornithidae Karkhu, 1988.Canadaga arctica Gen. nov. et Sp. nov. ValidHou LianhaiLate CretaceousMiddle Maastrichtian:A Hesperornithidae Marsh, 1872.Changchengornis hengdaoziensis Gen. nov. et Sp. nov. ValidJi Qiang
Luis M. Chiappe
Ji Shu’anEarly CretaceousChaomidianzi FormationA Confuciusornithidae Hou, Zhou, Gu et Zhang, 1995.Chaunoides antiquus Sp. nov. ValidHerculano M. F. de AlvarengaLate Oligocene or Early MioceneTremembé FormationAn Anhimidae.Chauvireria bulgarica Sp. nov. ValidBoevPleistoceneA Phasianidae.Confuciusornis dui Sp. nov. ValidHou Lianhai
Larry D. Martin
Zhou Zhonghe
Alan Feduccia
Zhang FuchengEarly CretaceousYixian FormationA Confuciusornithidae Hou, Zhou, Gu et Zhang, 1995.Dryolimnas augusti Sp. nov. ValidCécile Mourer-Chauviré
Roger Bour
Sonia Ribes
François MoutouHoloceneRéunionA Rallidae.Emberiza alcoveri Sp. nov. ValidJ. C. Rando
M. Lopez
B. SeguíLate Pleistocene-HoloceneCanary Islands, Tenerife:An Emberizidae.Eocuculus cherpinae Gen. nov. et Sp. nov.