Physiology and pharmacology Name Institution Year Peter Agre Johns Hopkins Bloomberg School of Public Health 2000 Richard W. Aldrich The University of Texas at Austin 2008 Susan G. Amara National Institutes of Health 2004 Clay M. Armstrong University of Pennsylvania 1987 Joseph Charles Aub (died 1973) Harvard University 1957 Kurt G. Beam University of Colorado School of Medicine 2012 Bruce P. Bean Harvard Medical School 2014 Joseph A. Beavo University of Washington School of Medicine 1996 Vann Bennett Duke University Medical Center 2010 Robert W. Berliner (died 2002) Yale School of Medicine 1968 Howard A. Bern (died 2012) University of California, Berkeley 1973 Robert M. Berne (died 2001) University of Virginia 1988 Michael J. Berridge (died 2020) Babraham Institute 1999 Karl H. Beyer Jr. (died 1996) Pennsylvania State University 1979 Francisco Bezanilla The University of Chicago 2006 Lutz Birnbaumer Pontifical Catholic University of Argentina 1995 James W. Black (died 2010) University of Dundee 1991 Barbara A. Block Stanford University 2023 John R. Brobeck (died 2009) University of Pennsylvania 1975 Bernard B. Brodie (died 1989) National Institutes of Health 1966 Chandler M. Brooks (died 1989) State University of New York 1975 Maurice B. Burg (died 2022) National Institutes of Health 1991 Arnold Burgen (died 2022) University of Cambridge 1987 John J. Burns (died 2007) Cornell University 1975 Michael D. Cahalan University of California, Irvine 2010 Nancy Carrasco Vanderbilt University School of Medicine 2015 William A. Catterall (died 2024) University of Washington 1989 W. Knox Chandler (died 2017) Yale University 1990 Julius H. Comroe Jr. (died 1984) University of California, San Francisco 1961 Allan H. Conney (died 2013) Rutgers University 1982 Erminio Costa (died 2009) University of Illinois at Chicago 1982 André Cournand (died 1988) Columbia University 1958 John W. Daly (died 2008) National Institutes of Health 1997 Horace W. Davenport (died 2005) University of Michigan 1974 Derek A. Denton (died 2022) University of Melbourne 1995 Peter N. Devreotes Johns Hopkins University School of Medicine 2005 Setsuro Ebashi (died 2006) National Institutes of Natural Sciences, Japan 1996 Robert H. Edwards University of California, San Francisco 2017 Vittorio Erspamer (died 1999) University of Rome 1990 John H. Exton (died 2022) Vanderbilt University 2001 Sérgio Henrique Ferreira (died 2016) University of São Paulo 2002 Robert Fettiplace University of Wisconsin-Madison 2021 Robert E. Forster (died 2021) University of Pennsylvania 1973 Clara Franzini-Armstrong University of Pennsylvania 1995 Robert F. Furchgott (died 2009) State University of New York Downstate Medical Center 1990 David L. Garbers (died 2006) University of Texas Southwestern Medical Center at Dallas 1993 Gerhard Giebisch (died 2020) Yale University 1984 Alfred G. Gilman (died 2015) University of Texas Southwestern Medical Center at Dallas 1985 Avram Goldstein (died 2012) Stanford University 1980 Louis Goodman (died 2000) University of Utah School of Medicine 1965 Ragnar Granit (died 1991) Karolinska Institutet 1968 James D. Hardy (died 1985) Yale University 1970 Steven C. Hebert (died 2008) Yale School of Medicine 2005 Peter Hegemann Humboldt University Berlin 2022 Yandell Henderson (died 1944) Yale University 1923 Donald W. Hilgemann UT Southwestern Medical Center 2021 Alan Hodgkin (died 1998) University of Cambridge 1974 Joseph F. Hoffman (died 2022) Yale University 1981 Jeffrey R. Holt Harvard Medical School 2025 A. James Hudspeth (died 2025) Rockefeller University 1991 Andrew Huxley (died 2012) University of Cambridge 1979 Louis J. Ignarro David Geffen School of Medicine at UCLA 1999 Holly A. Ingraham University of California, San Francisco 2021 Ehud Y. Isacoff University of California, Berkeley 2018 Laurinda A. Jaffe University of Connecticut Health Center 2021 Lily Y. Jan University of California, San Francisco School of Medicine 1995 Erik Jorgensen University of Utah 2022 David Julius University of California, San Francisco 2004 H. Ronald Kaback (died 2019) University of California, Los Angeles 1987 Arthur Karlin (died 2024) Columbia University 1999 Yoshito Kaziro (died 2011) Kyoto University 2000 Ernst Knobil (died 2000) University of Texas Health Science Center at Houston 1986 George B. Koelle (died 1997) University of Pennsylvania 1972 Edwin G. Krebs (died 2009) University of Washington 1973 Ching Kung University of Wisconsin-Madison 2011 Eugene M. Landis (died 1987) Harvard Medical School 1954 Ramón Latorre Universidad de Valparaiso 1999 Susan E. Leeman Boston University School of Medicine 1991 Greg Lemke Salk Institute for Biological Studies 2025 Richard S. Lewis Stanford University 2020 David P. Lloyd (died 1985) Rockefeller Institute 1953 Oliver H. Lowry (died 1996) Washington University School of Medicine in St. Louis 1964 David H. MacLennan (died 2020) University of Toronto 2001 Gail Mandel Oregon Health & Science University 2008 S. M. McCann (died 2007) Pennington Biomedical Research Center 1983 Randall T. Moon University of Washington 2015 Ferid Murad The George Washington University 1997 Philip Needleman (died 2024) Washington University in St. Louis 1987 Mark T. Nelson University of Vermont 2019 Baldomero M. Olivera University of Utah 2009 Krzysztof Palczewski University of California, Irvine 2022 John R. Pappenheimer (died 2007) Harvard University 1965 Charles R. Park (died 2016) Vanderbilt University 1980 Ardèm Patapoutian Scripps Research 2017 Eduardo A. Perozo The University of Chicago 2023 Donald W. Pfaff The Rockefeller University 1994 Tullio Pozzan (died 2022) University of Padua 2006 C. Ladd Prosser (died 2002) University of Illinois Urbana-Champaign 1974 Hermann Rahn (died 1990) University at Buffalo, The State University of New York 1968 Harald Reuter (died 2022) University of Bern 1997 Martin Rodbell (died 1998) National Institute of Environmental Health Sciences 1987 Juan Carlos Saéz Universidad de Valparaiso 2019 Charles H. Sawyer (died 2006) University of California, Los Angeles 1980 James A. Shannon (died 1994) National Institutes of Health 1965 Osamu Shimomura (died 2018) Marine Biological Laboratory 2013 Frederick J. Sigworth Yale School of Medicine 2016 Jens C. Skou (died 2018) University of Aarhus 1988 C. Richard Taylor (died 1995) Harvard University 1985 Chikashi Toyoshima The University of Tokyo 2005 Roger Y. Tsien (died 2016) University of California, San Diego 1998 Hans H. Ussing (died 2000) University of Copenhagen 1980 Wylie Vale (died 2012) Salk Institute for Biological Studies 1992 John Vane (died 2004) William Harvey Research Foundation 1983 Ewald R. Weibel (died 2019) University of Bern 1981 Gerald N. Wogan (died 2021) Massachusetts Institute of Technology 1977 Ernest M. Wright David Geffen School of Medicine at UCLA 2013 Nieng Yan Tsinghua University 2019 Masashi Yanagisawa UT Southwestern Medical Center 2003 Gary Yellen Harvard Medical School 2024