Name Institution Date Victor Ambros University of Massachusetts Medical School 2007 Angelika Amon (died 2020) MIT 2010 Kathryn Anderson (died 2020) Memorial Sloan-Kettering Cancer Center 2002 Thomas F. Anderson (died 1991) University of Pennsylvania 1964 Brenda Andrews University of Toronto 2020 Werner Arber University of Basel 1984 Ernest B. Babcock (died 1954) University of California, Berkeley 1946 Bruce Baker (died 2018) Stanford University 1993 Utpal Banerjee University of California, Los Angeles 2018 Rodolphe Barrangou North Carolina State University 2018 Bonnie Bartel Rice University 2016 George Beadle (died 1989) Stanford University 1944 Hugo J. Bellen Baylor College of Medicine 2020 Seymour Benzer (died 2007) California Institute of Technology 1961 Sue Biggins Fred Hutchinson Cancer Research Center 2015 Kerry Bloom University of North Carolina at Chapel Hill 2021 Jef D. Boeke New York University 2013 Nancy Bonini University of Pennsylvania 2012 Charles Boone University of Toronto 2024 David Botstein (died 2026) Princeton University 1981 Sydney Brenner (died 2019) Salk Institute for Biological Studies 1977 Royal Alexander Brink (died 1984) University of Wisconsin–Madison 1947 R. Daniel Camerini-Otero NIH 2025 Allan Campbell (died 2018) Stanford University 1971 John Carlson Yale University 2012 Marian Carlson Columbia University 2009 David Catcheside (died 1994) Australian National University 1974 L. L. Cavalli-Sforza (died 2018) Stanford University 1978 Martin Chalfie Columbia University 2004 Louise Chow University of Alabama at Birmingham 2012 Thomas W. Cline University of California, Berkeley 1996 Stanley N. Cohen Stanford University 1979 Victor Corces Emory University School of Medicine 2020 Elizabeth A. Craig University of Wisconsin–Madison 1998 James F. Crow (died 2012) University of Wisconsin–Madison 1961 Ronald W. Davis Stanford University 1983 Joseph DeRisi UC San Francisco 2016 August H. Doermann (died 1991) University of Washington 1975 William Dove University of Wisconsin–Madison 1998 Monica A. Driscoll Rutgers University 2023 Jay Dunlap Dartmouth College 2009 Susan K. Dutcher Washington University in St. Louis 2025 Harrison Echols (died 1993) University of California, Berkeley 1991 Robert Stuart Edgar (died 2016) University of California, Santa Cruz 2007 Evan E. Eichler University of Washington 2013 Sarah Elgin Washington University in St. Louis 2018 Scott Emmons Albert Einstein College of Medicine 2024 Ellis Englesberg University of California, Santa Barbara 1986 Stanley Fields University of Washington 2000 Gerald Fink Massachusetts Institute of Technology 1981 Andrew Fire Stanford University 2004 Maurice Fox (died 2020) Massachusetts Institute of Technology 1988 Antonio García-Bellido (died 2025) Autonomous University of Madrid 1987 Barry Ganetzky UW Madison 2006 Alan Garen (died 2022) Yale University 1971 Martin Gellert National Institutes of Health 1986 Michel A. J. Georges University of Liège 2013 Norman Giles (died 2006) University of Georgia 1966 N. Louise Glass University of California, Berkeley 2021 Michael M. Gottesman National Institutes of Health 2018 Susan Gottesman National Institutes of Health 1998 Daniel E. Gottschling Fred Hutchinson Cancer Research Center 2011 Melvin M. Green (died 2017) University of California, Davis 1980 Iva S. Greenwald Columbia University 2005 Shiv Grewal NIH 2014 Carol Gross University of California, San Francisco 1992 Alan D. Grossman MIT 2014 Christine Guthrie (died 2022) University of California, San Francisco 1993 James E. Haber Brandeis University 2010 Benjamin D. Hall (died 2019) University of Washington 2014 Jeffrey Hall Brandeis University 2003 Philip Hanawalt Stanford University 1989 Caryl Parker Haskins (died 2001) Carnegie Institution of Washington 1956 R. Scott Hawley (died 2025) Stowers Institute for Medical Research 2011 Ulrike Heberlein HHMI 2010 Donald Helinski University of California, San Diego 1980 Steven Henikoff Fred Hutchinson Cancer Research Center 2005 Tina Henkin Ohio State University 2012 Alfred Hershey (died 1997) Carnegie Institution of Washington 1958 Leonard Herzenberg (died 2013) Stanford University 1982 Philip Hieter University of British Columbia 2016 Alan Hinnebusch NIH 2015 Ann Hochschild Harvard Medical School 2024 Mark Hochstrasser Yale University 2025 Alexander Hollaender (died 1986) Associated Universities, Inc. 1957 Nancy Hopkins Massachusetts Institute of Technology 2004 Anita Hopper Ohio State University 2021 Norman Horowitz (geneticist) (died 2005) California Institute of Technology 1969 H. Robert Horvitz Massachusetts Institute of Technology 1991 David Housman Massachusetts Institute of Technology 1994 Malcolm Robert Irwin (died 1987) University of Wisconsin-Madison 1950 Harvey Itano (died 2010) University of California, San Diego 1979 Francois Jacob (died 2013) Institut Pasteur 1969 Patricia Jacobs University of Southampton 2009 Maria Jasin Memorial Sloan Kettering Cancer Center 2015 Nancy Jenkins Houston Methodist Research Institute 2008 Sue Jinks-Robertson Duke University School of Medicine 2019 Alexander D. Johnson UC San Francisco 2011 Mark Johnston University of Colorado School of Medicine 2022 A. Dale Kaiser (died 2010) Stanford University 1970 Berwind P. Kaufmann (died 1975) Cold Spring Harbor Laboratory 1952 Scott N. Keeney Memorial Sloan Kettering Cancer Center 2020 Cynthia Kenyon University of California, San Francisco 2003 Mary-Claire King University of Washington 2005 Hitoshi Kihara (died 1986) Kihara Institute for Biological Research 1958 Nancy Kleckner Harvard University 1993 Douglas Koshland UC Berkeley 2010 Mark Krasnow Stanford University School of Medicine 2019 Leonid Kruglyak University of California, Los Angeles 2023 Mitzi Kuroda Harvard University 2013 Sydney Kustu (died 2014) University of California, Berkeley 1993 Joshua Lederberg (died 2008) Rockefeller University 1957 Pedro León Azofefa Academia Nacional de Ciencias (Costa Rica) 2005 Leonard Lerman (died 2012) Massachusetts Institute of Technology 1986 Cyrus Levinthal (died 1990) Columbia University 1970 Edward B. Lewis (died 2004) California Institute of Technology 1968 Michael Lichten National Institutes of Health 2022 Susan Lindquist (died 2016) Massachusetts Institute of Technology 1997 Dan Lindsley (died 2018) University of California, San Diego 1974 David Lipman National Institutes of Health 2003 John T. Lis Cornell University 2015 Susan Lovett Brandeis University 2021 Victoria Lundblad Salk Institute for Biological Studies 2015 Salvador Luria (died 1991) Massachusetts Institute of Technology 1960 Mary F. Lyon (died 2014) Medical Research Council 1979 Boris Magasanik (died 2013) Massachusetts Institute of Technology 1969 Harmit Malik Fred Hutchinson Cancer Center 2019 Barbara McClintock (died 1992) Carnegie Institution of Washington 1944 Craig Mello University of Massachusetts Medical School 2005 M. S. Meselson Harvard University 1968 Charles W. Metz (died 1975) University of Pennsylvania 1948 Robert Metzenberg (died 2007) University of California, Los Angeles 1997 Barbara Meyer University of California, Berkeley 2000 Lois K. Miller (died 1999) University of Georgia 1997 Beatrice Mintz (died 2022) Fox Chase Cancer Center 1973 Danesh Moazed Harvard Medical School 2023 Hermann Joseph Muller (died 1967) University of Texas at Austin 1931 Andrew W. Murray Harvard University 2014 Terry Orr-Weaver MIT 2006 Howard Nash (died 2011) National Institutes of Health 1990 James V. Neel (died 2000) University of Michigan Medicine 1963 Elaine Ostrander National Human Genome Research Institute 2019 Ray David Owen (died 2014) California Institute of Technology 1966 David C. Page MIT 2005 Rafael Palacios Universidad Nacional Autonoma de Mexico 2006 Mary-Lou Pardue (died 2024) Massachusetts Institute of Technology 1983 David Perkins (died 2007) Stanford University 1981 Thomas Petes Duke University 1999 Guido Pontecorvo (died 1999) Imperial Cancer Research Fund 1983 John Preer Jr. (died 2016) Indiana University 1976 James Priess Fred Hutchinson Cancer Center 2017 John R. Pringle Stanford University School of Medicine 2017 Mark Ptashne Memorial Sloan-Kettering Cancer Center 1979 Theodore Puck (died 2005) University of Colorado Boulder 1960 Miroslav Radman Necker–Enfants Malades Hospital 2018 Marcus Morton Rhoades (died 1991) Indiana University 1946 Jasper Rine UC Berkeley 2008 Jeffrey Roberts Cornell University 1999 Shirleen Roeder Yale University 2009 Herschel L. Roman (died 1989) University of Washington 1970 John Roth University of California, Davis 1988 Lucia B. Rothman-Denes University of Chicago 2014 Rodney Rothstein Columbia University 2015 Gerald Rubin Howard Hughes Medical Institute 1987 Elizabeth S. Russell (died 2001) Jackson Laboratory 1972 Liane Russell (died 2019) Oak Ridge National Laboratory 1986 William L. Russell (died 2003) Oak Ridge National Laboratory 1973 Gary Ruvkun Harvard University 2008 Ruth Sager (died 1997) Dana–Farber Cancer Institute 1977 Margarita Salas (died 2019) Consejo Superior de Investigaciones Cientificas (CSIC) 2007 Matthew Scott Stanford University 1999 Julie Segre National Human Genome Research Institute 2022 Eric Selker University of Oregon 2012 Richard Setlow (died 2015) Brookhaven National Laboratory 1973 Geraldine Seydoux Johns Hopkins University 2016 Jay Shendure University of Washington 2022 Fred Sherman (died 2013) University of Rochester 1985 Obaid Siddiqi Tata Institute of Fundamental Research 2003 Melvin Simon California Institute of Technology 1985 Oliver Smithies (died 2017) University of North Carolina at Chapel Hill 1971 Franklin Stahl (died 2025) University of Oregon 1976 Peter Starlinger University of Cologne 1987 Kári Stefánsson University of Iceland 2019 George Streisinger (died 1984) University of Oregon 1975 Gisela Storz NIH 2012 F. William Studier Brookhaven National Laboratory 1992 Alfred Sturtevant (died 1970) California Institute of Technology 1930 Lorraine S. Symington Columbia University Irving Medical Center 2020 Jiazhen Tan (died 2008) Fudan University 1985 J. Herbert Taylor (died 1998) Florida State University 1977 Howard Martin Temin (died 1994) University of Wisconsin–Madison 1974 Jun-ichi Tomizawa (died 2017) National Institute of Genetics 1995 Bik Kwoon Tye Cornell University 2023 Anne Villeneuve (scientist) Stanford University School of Medicine 2017 Graham C. Walker MIT 2013 James D. Watson (died 2025) Cold Spring Harbor Laboratory 1962 Reed Wickner NIH 2000 Fred Winston Harvard University 2013 Evelyn Witkin (died 2023) Rutgers, The State University of New Jersey, New Brunswick 1977 Sewall Wright (died 1988) University of Wisconsin-Madison 1934 Élie Wollman (died 2008) Institut Pasteur 1991 Mariana Wolfner Cornell University 2019 William B. Wood III (died 2024) University of Colorado at Boulder 1972 Mitsuhiro Yanagida Okinawa Institute of Science and Technology 2012 Charles Yanofsky (died 2018) Stanford University 1966 Virginia Zakian Princeton University 2018 Phillip D. Zamore UMass Chan Medical School 2023 Norton Zinder (died 2012) Rockefeller University 1969