Name Institution Year C. David Allis (died 2023) The Rockefeller University 2005 Spyros Artavanis-Tsakonas Harvard Medical School 2023 Giuseppe Attardi (died 2008) California Institute of Technology 1984 Mary C. Beckerle Huntsman Cancer Institute 2021 Wolfgang Beermann (died 2000) Max Planck Institute for Biology 1975 Paul Berg (died 2023) Stanford University 1966 Eric Betzig University of California, Berkeley 2015 Wendy Bickmore University of Edinburgh 2025 Adrian Bird University of Edinburgh 2016 Max L. Birnstiel (died 2014) Intercell 1983 Mina J. Bissell Lawrence Berkeley National Laboratory 2010 Helen M. Blau Stanford University 2016 Günter Blobel (died 2018) Rockefeller University 1983 Gary G. Borisy The ADA Forsyth Institute 2009 Piet Borst Netherlands Cancer Institute 1991 Jean Brachet (died 1988) Free University of Brussels 1965 Robert W. Briggs (died 1983) Indiana University 1962 Ralph L. Brinster University of Pennsylvania 1987 Marianne Bronner California Institute of Technology 2015 Donald D. Brown (died 2023) Carnegie Institution of Washington 1973 Joan S. Brugge Harvard University 2001 Margaret Buckingham Institut Pasteur 2011 Gary Nathan Calkins (died 1943) Columbia University 1919 Lewis Clayton Cantley Dana-Farber Harvard Cancer Center 2001 Mario R. Capecchi University of Utah 1991 Howard Cedar Hebrew University of Jerusalem 2022 Constance L. Cepko Harvard University 2002 Pierre Chambon (died 2026) Institut de Genetique et de Biologie Moleculaire et Cellulaire (IGBMC) 1985 Don W. Cleveland University of California, San Diego 2006 Hans Clevers Roche Pharma Research & Early Development 2014 Stanley Cohen (died 2020) Vanderbilt University 1980 James E. Darnell Jr. Rockefeller University 1973 Eric H. Davidson (died 2015) California Institute of Technology 1985 Bernard D. Davis (died 1994) Harvard Medical School 1967 Igor B. Dawid (died 2024) National Institutes of Health 1981 Edward M. De Robertis David Geffen School of Medicine at UCLA 2013 Eugenia Del Pino Pontificia Universidad Católica del Ecuador 2006 Abby F. Dernburg University of California, Berkeley 2024 Claude Desplan New York University 2018 Chris Q. Doe University of Oregon 2017 David G. Drubin University of California, Berkeley 2022 Denis Duboule École Polytechnique Fédérale de Lausanne 2012 Peter H. Duesberg University of California, Berkeley 1986 Renato Dulbecco (died 2012) Salk Institute for Biological Studies 1961 Harry Eagle (died 1992) Albert Einstein College of Medicine 1963 William C. Earnshaw University of Edinburgh 2025 James D. Ebert (died 2001) Johns Hopkins University 1967 Bruce A. Edgar University of Utah 2025 Judith S. Eisen University of Oregon 2024 Scott D. Emr Cornell University 2007 John Franklin Enders (died 1985) Children’s Hospital Boston 1953 Anne Ephrussi European Molecular Biology Laboratory 2022 R. L. Erikson (died 2020) Harvard University 1983 Marilyn Gist Farquhar (died 2019) University of California, San Diego 1984 Don W. Fawcett (died 2009) Harvard University 1972 Gottfried S. Fraenkel (died 1984) University of Illinois Urbana-Champaign 1968 Elaine Fuchs Rockefeller University 1996 Margaret T. Fuller Stanford University 2008 Joseph G. Gall (died 2024) Carnegie Institution of Washington 1972 Walter J. Gehring (died 2014) University of Basel 1986 John C. Gerhart University of California, Berkeley 1990 Alfred L. Goldberg (died 2023) Harvard University 2015 Lawrence S. Goldstein University of California, San Diego 2020 Howard Green (died 2015) Harvard University 1978 Michael Green (died 2023) University of Massachusetts Medical School 2014 Clifford Grobstein (died 1998) University of California, San Diego 1966 Jerome Gross (died 2014) Harvard University 1974 John B. Gurdon (died 2025) University of Cambridge 1980 Karl Habel (died 1981) National Institute of Health 1968 W. A. Hagins (died 2012) National Institutes of Health 1979 Viktor Hamburger (died 2001) Washington University in St. Louis 1953 Richard M. Harland University of California, Berkeley 2014 Carl G. Hartman (died 1968) Carnegie Institution of Washington 1937 Leland H. Hartwell Fred Hutchinson Cancer Research Center 1987 Elizabeth D. Hay (died 2007) Harvard University 1984 Rebecca Heald University of California, Berkeley 2019 Edith Heard European Molecular Biology Laboratory 2021 Ira Herskowitz (died 2003) University of California, San Francisco 1986 Frederik L. Hisaw (died 1972) Harvard University 1947 Brigid L.M. Hogan Duke University 2005 David S. Hogness (died 2019) Stanford University 1976 Robert W. Holley (died 1993) Salk Institute for Biological Studies 1968 Johannes Holtfreter (died 1992) University of Rochester 1955 Rollin D. Hotchkiss (died 2004) Rockefeller University 1961 Tim Hunt Francis Crick Institute 1999 Anthony Hyman Max Planck Institute of Molecular Cell Biology and Genetics 2020 Richard O. Hynes (died 2026) Massachusetts Institute of Technology 1996 Shinya Inoué (died 2019) Marine Biological Laboratory 1993 Merkel Jacobs (died 1970) University of Pennsylvania 1939 Fotis C. Kafatos (died 2017) European Molecular Biology Laboratory 1982 Thomas C. Kaufman Indiana University 2008 Raymond E. Keller University of Virginia 2025 George Khoury (died 1987) National Cancer Institute 1987 Judith Kimble University of Wisconsin–Madison 1995 Charles B. Kimmel University of Oregon 2022 David M. Kingsley Stanford University 2011 Marc W. Kirschner Harvard University 1989 Aaron Klug (died 2018) Medical Research Council (UK) 1984 Thomas B. Kornberg University of California, San Francisco 2025 Robb Krumlauf Stowers Institute for Medical Research 2016 Nicole Le Douarin Académie des Sciences de l'Institut de France 1989 Philip Leder (died 2020) Harvard University 1979 Jeannie T. Lee Harvard University 2015 Ruth Lehmann New York University 2005 Maria Leptin European Research Council 2023 Michael S. Levine Princeton University 1998 Jennifer Lippincott-Schwartz National Institutes of Health 2008 Harvey F. Lodish Massachusetts Institute of Technology 1987 Richard M. Losick Harvard University 1992 David J. L. Luck (died 1998) Rockefeller University 1984 Anthony P. Mahowald University of Chicago 1994 Edward L. Mark (died 1946) Harvard University 1903 Gail R. Martin (died 2026) University of California, San Francisco 2002 Joan Massagué Memorial Sloan Kettering Cancer Center 2000 Iain Mattaj Human Technopole Foundation 2017 Satyajit (Jitu) Mayor National Centre for Biological Sciences 2015 Daniel Mazia (died 1996) Stanford University 1960 Clarence McClung (died 1946) University of Pennsylvania 1920 William J. McGinnis University of California, San Diego 2019 J. Richard McIntosh University of Colorado Boulder 1999 Steven L. McKnight University of Texas Southwestern Medical Center 1992 Andrew P. McMahon California Institute of Technology 2020 Peter Medawar (died 1987) Medical Research Council 1965 Douglas A. Melton Harvard University 1995 Oscar L. Miller Jr. (died 2012) University of Virginia 1978 Alfred Mirsky (died 1974) Rockefeller University 1954 Timothy J. Mitchison Harvard University 2014 Denise J. Montell University of California, Santa Barbara 2021 Melissa J. Moore University of Massachusetts Chan Medical School 2017 Ginés Morata Consejo Superior de Investigaciones Cientificas (CSIC) 2018 Aron Arthur Moscona (died 2009) University of Chicago 1977 Alex B. Novikoff (died 1987) Albert Einstein College of Medicine] 1974 Paul Nurse Francis Crick Institute 1995 Roeland Nusse Stanford University 2010 Christiane Nüsslein-Volhard Max Planck Institute for Developmental Biology 1990 Patrick H. O’Farrell University of California, San Francisco 2017 Eric N. Olson University of Texas Southwestern Medical Center 2000 Lelio Orci (died 2019) University of Geneva 1998 George E. Palade (died 2008) University of California, San Diego 1961 Arthur B. Pardee (died 2019) Harvard University 1968 Nipam H. Patel Marine Biological Laboratory 2024 Sheldon Penman (died 2021) Massachusetts Institute of Technology 1986 Norbert Perrimon Harvard University 2013 Robert Palese Perry (died 2013) Fox Chase Cancer Center 1977 Suzanne R. Pfeffer Stanford University 2024 Lennart Philipson (died 2011) Karolinska Institute 1992 Keith R. Porter (died 1997) University of Pennsylvania 1964 Olivier Pourquié Brigham and Women's Hospital 2020 David M. Prescott (died 2011) University of Colorado at Boulder 1974 Martin C. Raff University College London 2003 Anjana Rao La Jolla Institute for Immunology 2008 Hans Ris (died 2004) University of Wisconsin-Madison 1974 Elizabeth J. Robertson University of Oxford 2021 Robert G. Roeder Rockefeller University 1988 Michael Rosbash Brandeis University 2003 Janet Rossant University of Toronto 2008 James E. Rothman Columbia University 1993 Francis H. Ruddle (died 2013) Yale University 1976 Joan V. Ruderman Harvard University 1998 David D. Sabatini New York University 1985 David M. Sabatini Massachusetts Institute of Technology 2016 Edward D. Salmon University of North Carolina at Chapel Hill 2021 Alejandro Sánchez Alvarado Stowers Institute for Medical Research 2018 Gordon H. Sato (died 2017) Ministry of Fisheries of Eritrea 1984 John W. Saunders Jr. (died 2015) State University of New York at Albany 2006 Randy Wayne Schekman University of California, Berkeley 1992 Alexander F. Schier University of Basel 2020 Sandra L. Schmid Chan Zuckerberg Biohub 2020 Karl Patterson Schmidt (died 1957) Field Museum 1956 Howard A. Schneiderman (died 1990) Monsanto Company 1975 Gertrud M. Schüpbach Princeton University 2005 Lucy Shapiro Stanford University 1994 Phillip A. Sharp Massachusetts Institute of Technology 1983 Aaron J. Shatkin (died 2012) Rutgers University 1981 Philip Siekevitz (died 2009) Rockefeller University 1975 Pamela A. Silver Harvard University 2023 Kai Simons Max Planck Institute for Molecular Cell Biology and Genetics 1997 Robert H. Singer Albert Einstein College of Medicine 2013 S. J. Singer (died 2017) University of California, San Diego 1969 Allan C. Spradling Carnegie Institution for Science 1989 James A. Spudich Stanford University 1991 Paul W. Sternberg California Institute of Technology 2009 Charles Stockard (died 1939) Cornell Medical College 1922 William Straus Jr. (died 1981) Johns Hopkins University 1962 Susan Strome University of California, Santa Cruz 2019 Gary Struhl Columbia University 2008 Kevin Struhl Harvard University 2010 Hewson Swift (died 2004) The University of Chicago 1971 Clifford J. Tabin Harvard University 2007 Masatoshi Takeichi RIKEN 2007 Elly M. Tanaka Institute of Molecular Biotechnology 2023 Edwin W. Taylor Northwestern University 2001 Julie A. Theriot University of Washington 2021 Irma Thesleff University of Helsinki 2017 Shirley M. Tilghman Princeton University 1996 Lewis G. Tilney University of Pennsylvania 1998 Robert Tjian University of California, Berkeley 1991 Ronald D. Vale University of California, San Francisco 2001 K. VijayRaghavan National Centre for Biological Sciences 2014 Salome G. Waelsch (died 2007) Yeshiva University 1979 Clare M. Waterman National Institutes of Health 2018 Fiona M. Watt EMBO 2019 Paul A. Weiss (died 1989) The University of Chicago 1947 Eric F. Wieschaus Princeton University 1994 Vincent Wigglesworth (died 1994) University of Cambridge 1971 Carroll M. Williams (died 1991) Harvard University 1960 Simeon Burt Wolbach (died 1954) Harvard Medical School 1938 Jerry L. Workman Stowers Institute for Medical Research 2025 Joanna K. Wysocka Stanford University 2024 Keith R. Yamamoto University of California, San Francisco 1990 Yukiko Yamashita Whitehead Institute for Biomedical Research 2025 Michael W. Young The Rockefeller University 2007 Richard A. Young Massachusetts Institute of Technology 2012 Junying Yuan Chinese Academy of Sciences 2017 Kenneth Zaret University of Pennsylvania 2023 Raymond E. Zirkle (died 1988) The University of Chicago 1959