Name Institution Year Robert H. Abeles (died 2000) Brandeis University 1976 John Abelson California Institute of Technology 1985 Julius Adler (died 2024) University of Wisconsin–Madison 1978 Paul Ahlquist University of Wisconsin–Madison 1993 Natalie G. Ahn University of Colorado Boulder 2018 Bruce Alberts University of California, San Francisco 1981 Jorge E. Allende University of Chile 2001 Sidney Altman (died 2022) Yale University 1990 Bruce Ames (died 2024) University of California, Berkeley 1972 Christian B. Anfinsen (died 1995) Johns Hopkins University School of Medicine 1963 Daniel I. Arnon (died 1994) University of California, Berkeley 1961 Gilbert Ashwell (died 2014) National Institutes of Health 1979 Tania Baker Massachusetts Institute of Technology 2007 Clinton Ballou (died 2021) University of California, Berkeley 1975 Horace Barker (died 2000) University of California, Berkeley 1953 David P. Bartel Massachusetts Institute of Technology 2011 Jacqueline Barton California Institute of Technology 2002 Brenda Bass University of Utah School of Medicine 2015 Philip Beachy Stanford University School of Medicine 2002 Lorena S. Beese Duke University Medical Center 2009 Marlene Belfort New York State Department of Health 1999 Stephen P. Bell Massachusetts Institute of Technology 2017 James M. Berger Johns Hopkins University School of Medicine 2013 Sune Bergström (died 2004) Karolinska Institutet 1973 Klaus Biemann (died 2016) Massachusetts Institute of Technology 1993 Pamela Björkman California Institute of Technology 2001 Elizabeth Blackburn University of California, San Francisco 1993 Konrad E. Bloch (died 2000) Harvard University 1956 Elkan Blout (died 2006) Harvard University 1969 Squire J. Booker University of Pennsylvania 2019 Michael Botchan University of California, Berkeley 2008 Herbert Boyer University of California, San Francisco 1985 Paul D. Boyer (died 2018) University of California, Los Angeles 1970 Roscoe Brady (died 2016) National Institutes of Health 1975 Alexander E. Braunstein (died 1986) USSR Academy of Medical Sciences 1974 Ronald Breaker Yale University 2013 Joan B. Broderick Montana State University 2022 Patrick O. Brown Impossible Foods 2002 John Buchanan (died 2007) Massachusetts Institute of Technology 1962 Walter B. Cannon (died 1945) Harvard Medical School 1914 Giulio Cantoni (died 2005) National Institutes of Health 1983 John Carbon University of California, Santa Barbara 1986 Dana Carroll University of Utah School of Medicine 2017 H. E. Carter (died 2007) University of Arizona 1953 Thomas R. Cech University of Colorado Boulder 1987 Michael Chamberlin (died 2025) University of California, Berkeley 1986 Britton Chance (died 2010) University of Pennsylvania 1954 Erwin Chargaff (died 2002) Columbia University 1965 Aaron Ciechanover Technion-Israel Institute of Technology 2007 W. Wallace Cleland (died 2013) University of Wisconsin–Madison 1985 Philip Cohen University of Dundee 2008 Seymour Cohen (died 2018) Marine Biological Laboratory 1967 Mildred Cohn (died 2009) University of Pennsylvania 1971 Kathleen Collins University of California, Berkeley 2025 Sidney Colowick (died 1985) Vanderbilt University 1972 Joan W. Conaway UT Southwestern Medical Center 2020 Elena Conti Max Planck Institute of Biochemistry 2025 Minor Coon (died 2018) University of Michigan 1983 Carl F. Cori (died 1984) Harvard University 1940 Nicholas R. Cozzarelli (died 2006) University of California, Berkeley 1989 Nancy Craig Johns Hopkins University 2010 Patrick Cramer Max Planck Institute for Multidisciplinary Sciences 2020 Benjamin Cravatt III Scripps Research 2014 James Dahlberg University of Wisconsin–Madison 1996 Seth Darst Rockefeller University 2008 Earl Davie (died 2020) University of Washington 1980 Christian de Duve (died 2013) Université catholique de Louvain 1975 Hector DeLuca University of Wisconsin–Madison 1979 Peter Dervan California Institute of Technology 1986 Raymond Deshaies Amgen 2016 John F.X. Diffley Francis Crick Institute 2020 Zacharias Dische (died 1988) Columbia University 1976 Jack Dixon University of California, San Diego 2000 Edward A. Doisy (died 1986) Saint Louis University 1938 Russell Doolittle (died 2019) University of California, San Diego 1984 Paul M. Doty (died 2011) Harvard University 1957 Jennifer Doudna University of California, Berkeley 2002 Catherine L. Drennan Massachusetts Institute of Technology 2023 Gideon Dreyfuss University of Pennsylvania 2012 Gerald Edelman (died 2014) The Scripps Research Institute 1969 Isidore S. Edelman (died 2004) Columbia University 1973 Mary Edmonds (died 2005) University of Pittsburgh 1991 John T. Edsall (died 2002) Harvard University 1951 Michael B. Elowitz California Institute of Technology 2022 Ronald Estabrook (died 2013) University of Texas Southwestern Medical Center at Dallas 1979 Gerald D. Fasman (died 2003) Brandeis University 1994 Gary Felsenfeld (died 2024) National Institutes of Health 1976 Edmond Fischer (died 2021) University of Washington 1973 Heinz Fraenkel-Conrat (died 1999) University of California, Berkeley 1974 Perry A. Frey University of Wisconsin–Madison 1998 Irwin Fridovich (died 2019) Duke University 1981 Morris E. Friedkin (died 2002) University of California, San Diego 1978 Joseph S. Fruton (died 2007) Yale University 1952 Judith Frydman Stanford University 2021 Mariano A. Garcia-Blanco University of Virginia 2025 E. Peter Geiduschek (died 2022) University of California, San Diego 1974 Quentin Gibson (died 2011) Rice University 1982 Lila M. Gierasch University of Massachusetts Amherst 2019 Alexander Glazer (died 2021) University of California System 2001 Sam Granick (died 1977) Rockefeller Institute for Medical Research 1965 David Green (died 1983) University of Wisconsin-Madison 1962 Rachel Green Johns Hopkins University 2012 Carol Greider University of California, Santa Cruz 2003 Jack D. Griffith University of North Carolina at Chapel Hill 2018 Marianne Grunberg-Manago (died 2013) Institut de Biologie Physico-Chimique 1982 Michael Grunstein (died 2024) University of California, Los Angeles 2008 Irwin Gunsalus (died 2008) University of Illinois at Urbana–Champaign 1965 Lowell Hager (died 2014) University of Illinois at Urbana–Champaign 1995 Sen-itiroh Hakomori Pacific Northwest Research Institute 2000 Heidi Hamm Vanderbilt University 2025 Gregory Hannon (died 2026) University of Cambridge 2012 J. Wade Harper Harvard Medical School 2023 William F. Harrington (died 1992) Johns Hopkins University School of Medicine 1976 F. Ulrich Hartl Max Planck Institute of Biochemistry 2011 Felix Haurowitz (died 1987) Indiana University 1975 Ari Helenius ETH Zurich 2009 Richard Henderson Medical Research Council 1998 Wayne Hendrickson Columbia University 1993 Leon A. Heppel (died 2010) Cornell University 1970 Dan Herschlag Stanford University 2018 Avram Hershko Technion Israel Institute of Technology 2003 George Hess (died 2015) Cornell University 1987 Robert L. Hill (died 2012) Duke University 1975 Mahlon Hoagland (died 2009) Worcester Foundation for Biomedical Research 1984 Klaus Hofmann (died 1995) University of Pittsburgh 1963 B. L. Horecker (died 2010) Cornell University 1961 Arthur Horwich Yale University 2003 Robert Huber Max Planck Institute for Biochemistry 1995 Jerard Hurwitz (died 2019) Memorial Sloan-Kettering Cancer Center 1974 Clyde A. Hutchison III (died 2025) J. Craig Venter Institute 1995 Masayori Inouye Rutgers University 2019 William Jencks (died 2007) Brandeis University 1971 Mary Ellen Jones (died 1996) University of North Carolina, School of Medicine 1984 Leemor Joshua-Tor Cold Spring Harbor Laboratory 2017 Gerald Joyce Salk Institute for Biological Studies 2001 Elvin Kabat (died 2000) Columbia University College of Physicians and Surgeons 1966 James Kadonaga University of California, San Diego 2022 Paul Kaesberg (died 2010) University of Wisconsin–Madison 1991 Herman Kalckar (died 1991) Harvard University 1959 Martin Kamen (died 2002) University of California, San Diego 1962 Nathan Kaplan (died 1986) University of California, San Diego 1970 Seymour Kaufman (died 2009) National Institutes of Health 1986 Thomas J. Kelly Johns Hopkins University 1992 John Kendrew (died 1997) University of Oxford 1972 Eugene P. Kennedy (died 2011) Harvard University 1964 Har Gobind Khorana (died 2011) Massachusetts Institute of Technology 1966 Peter Kim Stanford University 1997 V. Narry Kim Seoul National University 2014 Robert Kingston Harvard University 2016 Rachel Klevit University of Washington 2021 Judith Klinman University of California, Berkeley 1994 Irving Klotz (died 2005) Northwestern University 1970 Jeremy Knowles (died 2008) Harvard University 1988 Edward Korn National Institutes of Health 1990 Arthur Kornberg (died 2007) Stanford University 1957 Roger Kornberg Stanford University 1993 Alberto Kornblihtt University of Buenos Aires 2011 Daniel E. Koshland, Jr. (died 2007) University of California, Berkeley 1966 Stephen Kowalczykowski University of California, Davis 2007 Adrian Krainer Cold Spring Harbor Laboratory 2020 Lester O. Krampitz (died 1993) Case Western Reserve University 1978 Joseph Kraut (died 2012) University of California, San Diego 1988 Thomas A. Kunkel National Institute of Environmental Health Sciences 2024 Alan Lambowitz University of Texas at Austin 2004 Rebecca Lancefield (died 1981) Rockefeller University 1970 Arthur Landy Brown University 1999 M. Daniel Lane (died 2014) Johns Hopkins University 1987 Henry Lardy (died 2010) University of Wisconsin–Madison 1958 I. Robert Lehman (died 2026) Stanford University 1977 Albert Lehninger (died 1986) Johns Hopkins University School of Medicine 1956 Luis F. Leloir (died 1987) Instituto de Investigaciones Bioquimicas of Argentina 1960 William Lennarz (died 2021) State University of New York at Stony Brook 1989 Choh Hao Li (died 1987) University of California, San Francisco 1973 Christopher D. Lima Memorial Sloan Kettering Cancer Center 2020 Tomas Lindahl Francis Crick Institute 2018 Fritz Lipmann (died 1986) Rockefeller University 1950 Laszlo Lorand (died 2018) Northwestern University 1987 George Lorimer University of Maryland, College Park 1997 Martha Ludwig (died 2006) University of Michigan 2003 Feodor Lynen (died 1979) Max Planck Institute of Biochemistry 1962 James Manley Columbia University 2011 Matthias Mann Max Planck Institute of Biochemistry 2025 Lynne E. Maquat University of Rochester 2011 Emanuel Margoliash (died 2008) Northwestern University 1975 Michael Marletta University of California, Berkeley 2006 Vincent Massey (died 2002) University of Michigan School of Medicine 1995 Rowena Green Matthews University of Michigan 2002 William D. McElroy (died 1999) University of California, San Diego 1963 Alton Meister (died 1995) Cornell University 1969 Robert Bruce Merrifield (died 2006) Rockefeller University 1972 Karl Meyer (died 1990) Columbia University 1967 Stanley Miller (died 2007) University of California, San Diego 1973 Peter D. Mitchell (died 1992) Glynn Research Institute 1977 Kiyoshi Mizuuchi National Institutes of Health 1994 Paul Modrich Duke University 1993 Tom Muir Princeton University 2024 Geeta Narlikar University of California, San Francisco 2021 Daniel Nathans (died 1999) Johns Hopkins University School of Medicine 1979 Elizabeth Neufeld University of California, Los Angeles 1977 Hans Neurath (died 2002) University of Washington 1961 Marshall Warren Nirenberg (died 2010) National Institutes of Health 1967 Yasutomi Nishizuka (died 2004) Kobe University 1988 Harry Noller