List of Chthoniidae species

This is a list of the described species of the pseudoscorpion family Chthoniidae. The data is taken from Joel Hallan's Biology Catalog.

Afrochthonius Beier, 1930 Afrochthonius brincki Beier, 1955 — Lesotho Afrochthonius ceylonicus Beier, 1973 — Sri Lanka Afrochthonius godfreyi (Ellingsen, 1912) — Southern Africa Afrochthonius inaequalis Beier, 1958 — Southern Africa Afrochthonius natalensis Beier, 1931 — Southern Africa Afrochthonius reductus Beier, 1973 — Sri Lanka Afrochthonius similis Beier, 1930 — Namibia

Allochthonius J.C. Chamberlin, 1929 Allochthonius borealis Sato, 1984 — Japan Allochthonius buanensis W.K.Lee, 1982 — Korea Allochthonius kinkaiensis Sakayori, 2002 — Japan Allochthonius montanus Sakayori, 2000 — Honshu, Tochigi, Mount Kohshin, Japan Allochthonius opticus (Ellingsen, 1907) — Japan Allochthonius opticus opticus (Ellingsen, 1907) — Japan Allochthonius opticus coreanus Morikawa, 1970 — Korea Allochthonius opticus troglophilus Morikawa, 1956 — Japan Allochthonius shintoisticus J.C. Chamberlin, 1929 — Japan Allochthonius tamurai Sakayori, 1999 — Japan Allochthonius biocularis (Morikawa, 1956) — Japan Allochthonius ishikawai (Morikawa, 1954) — Japan Allochthonius ishikawai ishikawai (Morikawa, 1954) — Japan Allochthonius ishikawai deciclavatus (Morikawa, 1956) — Japan Allochthonius ishikawai kyushuensis (Morikawa, 1960) — Japan Allochthonius ishikawai shiragatakiensis (Morikawa, 1954) — Japan Allochthonius ishikawai uenoi (Morikawa, 1956) — Japan Allochthonius uyamadensis (Morikawa, 1954) — Japan

Aphrastochthonius J.C. Chamberlin, 1962 Aphrastochthonius alteriae Muchmore, 1977 — Mexico Aphrastochthonius cubanus Dumitresco & Orghidan, 1977 — Cuba Aphrastochthonius grubbsi Muchmore, 1984 — California Aphrastochthonius major Muchmore, 1973 — Mexico Aphrastochthonius pachysetus Muchmore, 1976 — New Mexico Aphrastochthonius palmitensis Muchmore, 1986 — Mexico Aphrastochthonius parvus Muchmore, 1972 — Mexico Aphrastochthonius patei Muchmore, 1982 — Mexico Aphrastochthonius pecki Muchmore, 1968 — Alabama Aphrastochthonius russelli Muchmore, 1972 — Mexico Aphrastochthonius similis Muchmore, 1984 — California Aphrastochthonius tenax J.C. Chamberlin, 1962 Aphrastochthonius verapazanus Muchmore, 1972 — Guatemala

Apochthonius J.C. Chamberlin, 1929 Apochthonius coecus (Packard, 1884) — Virginia Apochthonius colecampi Muchmore, 1967 — Missouri Apochthonius diabolus Muchmore, 1967 — Arkansas Apochthonius forbesi Benedict, 1979 — Oregon Apochthonius grubbsi Muchmore, 1980 — California Apochthonius hobbsi Muchmore, 1994 — Ohio Apochthonius holsingeri Muchmore, 1967 — Virginia Apochthonius hypogeus Muchmore, 1976 — Virginia Apochthonius indianensis Muchmore, 1967 — Indiana Apochthonius intermedius J.C. Chamberlin, 1929 — Washington Apochthonius irwini R.O. Schuster, 1966 — California Apochthonius knowltoni Muchmore, 1980 — Wyoming Apochthonius magnanimus Hoff, 1956 — New Mexico Apochthonius malheuri Benedict & Malcolm, 1973 — Oregon Apochthonius maximus R.O. Schuster, 1966 — California Apochthonius minimus R.O. Schuster, 1966 — British Columbia, Oregon, Washington Apochthonius minor Muchmore, 1976 — Georgia Apochthonius moestus (Banks, 1891) — Eastern U.S. Apochthonius mysterius Muchmore, 1976 — Missouri Apochthonius occidentalis J.C. Chamberlin, 1929 — Oregon Apochthonius paucispinosus Muchmore, 1967 — West Virginia Apochthonius russelli Muchmore, 1976 — Alabama Apochthonius titanicus Muchmore, 1976 — Arkansas Apochthonius typhlus Muchmore, 1967 — Missouri

Austrochthonius J.C. Chamberlin, 1929 Austrochthonius argentinae Hoff, 1950 — Argentina, Brazil Austrochthonius australis Hoff, 1951 — Australia Austrochthonius bolivianus Beier, 1930 — Bolivia Austrochthonius cavicola Beier, 1968 — Australia Austrochthonius chilensis (J.C. Chamberlin, 1923) Austrochthonius chilensis chilensis (J.C. Chamberlin, 1923) — Chile, Argentina Austrochthonius chilensis magalhanicus Beier, 1964 — Chile Austrochthonius chilensis transversus Beier, 1964 — Argentina Austrochthonius easti Harvey, 1991 — western Australia Austrochthonius iguazuensis Vitali-di Castri, 1975 — Brazil Austrochthonius insularis Vitali-di Castri, 1968 — Crozet Islands Austrochthonius mordax Beier, 1967 — New Zealand Austrochthonius paraguayensis Vitali-di Castri, 1975 — Paraguay Austrochthonius parvus (Mello-Leitão, 1939) — Argentina Austrochthonius persimilis Beier, 1930 — Chile Austrochthonius rapax Beier, 1976 — New Zealand Austrochthonius semiserratus Beier, 1930 — Chile Austrochthonius tullgerni (Beier, 1931) — Southern Africa Austrochthonius zealandicus Beier, 1966 — New Zealand Austrochthonius zealandicus zealandicus Beier, 1966 — New Zealand Austrochthonius zealandicus obscurus Beier, 1966 — New Zealand

Caribchthonius Muchmore, 1976 Caribchthonius butleri Muchmore, 1976 — Virgin Islands Caribchthonius orthodentatus Muchmore, 1976 — Belize

Centrochthonius Beier, 1931 Centrochthonius koslovi (Redikorzev, 1918) — Tibet, India, Nepal Centrochthonius schnitnikovi (Redikorzev, 1934) — Russia Centrochthonius sichuanensis Schawaller, 1995 — Sichuan

Chiliochthonius Vitali-di Castri, 1975 Chiliochthonius centralis Vitali-di Castri, 1975 — Chile Chiliochthonius montanus Vitali-di Castri, 1975 — Chile Chiliochthonius usuriensis Beier, 1979 — Russia (?)

Chthonius C.L. Koch, 1843 Chthonius absoloni Beier, 1938 — Yugoslavia Chthonius agazzii Beier, 1966 — Italy Chthonius alpicola Beier, 1951 — Austria, Italy Chthonius apollinis Mahnert, 1978 — Greece Chthonius azerbaidzhanus Schawaller & Dashdamirov, 1988 — Azerbaijan Chthonius baccettii Callaini, 1980 — Italy Chthonius balazuci Vachon, 1963 — France Chthonius bogovinae Curcic, 1972 — Yugoslavia Chthonius bogovinae bogovinae Curcic, 1972 — Yugoslavia Chthonius bogovinae latidentatus Curcic, 1972 — Yugoslavia Chthonius brandmayri Callaini, 1986 — Yugoslavia Chthonius caprai Gardini, 1977 — Italy Chthonius caoduroi Callaini, 1987 — Italy Chthonius cavernarum Ellingsen, 1909 — Romania, Yugoslavia Chthonius cavophilus Hadzi, 1939 — Bulgaria Chthonius cebenicus Leclerc, 1981 — France Chthonius cephalotes (Simon, 1875) — France Chthonius chamberlini (Leclerc, 1983) — Algeria, France Chthonius comottii Inzaghi, 1987 — Italy Chthonius cryptus J.C. Chamberlin, 1962 — Greece Chthonius dacnodes Navás, 1918 — Spain, Azores, Italy Chthonius dalmatinus Hadzi, 1930 — Yugoslavia Chthonius densedentatus Beier, 1938 — Italy, Yugoslavia Chthonius diophthalmus Daday, 1888 — eastern Europe Chthonius doderoi Beier, 1930 — France, Italy Chthonius doderoi doderoi Beier, 1930 — France, Italy Chthonius doderoi horridus Beier, 1934 — Italy Chthonius ellingseni Beier, 1928 — Italy, Yugoslavia Chthonius elongatus Lazzeroni, 1970 — Italy Chthonius euganeus Gardini, 1991 — Italy Chthonius exarmatus Beier, 1939 — Yugoslavia Chthonius graecus Beier, 1963 — Greece Chthonius guglielmii Callaini, 1986 — Italy Chthonius halberti Kew, 1916 — France, Great Britain Chthonius herbarii Mahnert, 1980 — Crete Chthonius heterodactylus Tomosvary, 1882 — central Europe Chthonius heutaultae Leclerc, 1981 — France Chthonius hungaricus Mahnert, 1981 — Hungary Chthonius ilvensis Beier, 1963 — Italy Chthonius imperator Mahnert, 1978 — Greece Chthonius ischnocheles (Hermann, 1804) — Europe, U.S. Chthonius ischnocheles ischnocheles (Hermann, 1804) — Europe, U.S. Chthonius ischnocheles reductus Beier, 1939 — southern Europe Chthonius ischnocheles ruffoi Caporiacco, 1951 — Italy Chthonius ischnocheles stammeri Beier, 1942 — Italy Chthonius ischnocheloides Beier, 1974 — Italy Chthonius italicus Beier, 1930 — Italy Chthonius iugoslavicus Curcic, 1972 — Yugoslavia Chthonius jalzici Curcic, 1988 — Yugoslavia Chthonius jonicus Beier, 1931 — southern Europe Chthonius jugorum Beier, 1952 — Italy Chthonius karamanianus Hadzi, 1937 — Yugoslavia Chthonius lanzai Caporiacco, 1947 — Italy Chthonius lanzai lanzai Caporiacco, 1947 — Italy Chthonius lanzai vannii Callaini, 1986 — Italy Chthonius leoi (Callaini, 1988) — Sardinia Chthonius leruthi Beier, 1939 — Romania Chthonius lesnik Curcic, 1994 — Yugoslavia Chthonius lessiniensis Schawaller, 1982 — Italy Chthonius ligusticus Beier, 1930 — Italy Chthonius lindbergi Beier, 1956 — Crete Chthonius litoralis Hadzi, 1933 — Italy, Yugoslavia Chthonius lucifugus Mahnert, 1977 — France, Spain Chthonius macedonicus Curcic, 1972 — Yugoslavia Chthonius magnificus Beier, 1938 — Yugoslavia Chthonius malatestai Callaini, 1980 — Italy Chthonius mauritanicus (Callaini, 1988) — Morocco Chthonius mayi Heurtault-Rossi, 1968 — France Chthonius mazaurici Leclerc, 1981 — France Chthonius mazaurici mazaurici Leclerc, 1981 — France Chthonius mazaurici coironi Leclerc, 1981 — France † Chthonius mengei Beier, 1937 — fossil: Baltic amber Chthonius microphthalmus Simon, 1879 — France, Spain Chthonius mingazzinii Callaini, 1991 — Italy Chthonius minotaurus Henderickx, 1997 — Kournas Cave, Crete Chthonius monicae Boghean, 1989 — Romania Chthonius motasi Dumitresco & Orghidan, 1964 — Romania Chthonius multidentatus Beier, 1963 — Sicily Chthonius occultus Beier, 1939 — Yugoslavia Chthonius orthodactyloides Beier, 1967 — Turkey Chthonius orthodactylus (Leach, 1817) — Europe Chthonius pacificus Muchmore, 1968 — California Chthonius paganus (Hoff, 1961) — Colorado Chthonius paludis (J.C. Chamberlin, 1929) — southern U.S. Chthonius paolettii Beier, 1974 — Italy Chthonius persimilis Beier, 1939 — Yugoslavia Chthonius petrochilosi Heurtault, 1972 — Greece Chthonius pivai Gardini, 1991 — Italy Chthonius ponticoides Mahnert, 1975 — Greece Chthonius ponticus Beier, 1965 — Turkey Chthonius porevidi Curcic, Makarov & Lucic, 1998 — Krivosije, Knezlaz Pecina Cave, Montenegro † Chthonius pristinus Schawaller, 1978 — fossil: Baltic amber Chthonius prove Curcic, Dimitrijevic & Makarov, 1997 — Gornji Morinj, Serbia, Montenegro Chthonius pusillus Beier, 1947 — Austria Chthonius pygmaeus Beier, 1934 — central Europe Chthonius pygmaeus pygmaeus Beier, 1934 — central Europe Chthonius pygmaeus carinthiacus Beier, 1951 — central Europe Chthonius radjai Curcic, 1988 — Yugoslavia Chthonius raridentatus Hadzi, 1930 — Yugoslavia Chthonius ressli Beier, 1956 — Austria, France, Italy Chthonius satapliaensis Schawaller & Dashdamirov, 1988 — Russia Chthonius sestasi Mahnert, 1980 — Greece Chthonius shelkovnikovi Redikorzev, 1930 — western Asia Chthonius shulovi Beier, 1963 — Israel Chthonius stevanovici Curcic, 1986 — Yugoslavia Chthonius strinatii Mahnert, 1975 — Greece Chthonius submontanus Beier, 1963 — central Europe Chthonius subterraneus Beier, 1931 — Hungary, Yugoslavia Chthonius subterraneus subterraneus Beier, 1931 — Hungary, Yugoslavia Chthonius subterraneus meuseli Beier, 1939 — Yugoslavia Chthonius tadzhikistanicus Dashdamirov & Schawaller, 1992 — Tajikistan Chthonius tenuis L. Koch, 1873 — Europe, Algeria Chthonius thessalus Mahnert, 1980 — Greece Chthonius trebinjensis Beier, 1938 — Yugoslavia Chthonius troglobius Hadzi, 1937 — Yugoslavia Chthonius troglodites Redikorzev, 1928 — Bulgaria Chthonius tzanoudakisi Mahnert, 1975 — Greece Chthonius aegatensis (Callaini, 1989) — Sicilly Chthonius aguileraorum Carabajal Marquez, Garcia Carrillo & Rodriguez Fernandez, 2000 — Cadiz, Villaluenga del Rosario, Cueva de la Hiedra, Spain Chthonius amatei Carabajal Marquez, Garcia Carrillo & Rodriguez Fernandez, 2001 — Spain Chthonius anatolicus (Beier, 1969) — Iran, Turkey Chthonius anophthalmus (Ellingsen, 1908) — Algeria Chthonius apulicus (Beier, 1958) — Italy Chthonius asturiensis (Beier, 1955) — Spain Chthonius atlantis (Mahnert, 1980) — Morocco Chthonius balearicus (Mahnert, 1977) — Balearic Islands Chthonius bartolii (Gardini, 1976) — Italy Chthonius bauneensis (Callaini, 1983) — Sardinia