Argiles et Grès à Reptiles Formation (meaning Reptile Clay and Sandstone) also known as the Argiles Rutilantes FormationTabuce et al., 2013 is an early MaastrichtianWeishampel et al., 2004, pp.588-593 French geologic formation in the département of Var preserving the remains of several types of dinosaurs and other extinct organisms.Argiles et Grès à Reptiles Formation at Fossilworks.org
Fossil content An abelisaurid similar to Arcovenator is known from Département des Bouches-du-Rhone, while enantiornithean and indeterminate avialan remains are known from Département du Var.
Dinosaurs Ornithischians Ornithischians of the Argiles et Grès à Reptiles Formation Genus Species Location Stratigraphic position Material Notes Images Matheronodon M. provincialis Velaux-La Bastide Neuve Lower A single maxilla and associated teeth A rhabdodontid ornithopod Obelignathus O. septimanicus Montouliers, Saint-Chinian commune Lower An isolated right dentary, missing the anterior (front) part. A rhabdodontomorph iguanodontian RhabdodonChanthasit, P. (2010). The ornithopod dinosaur Rhabdodon from the Late Cretaceous of France : anatomy, systematics and paleobiology. Http Www Theses Fr. R. priscus La Nerthe Upper A left dentary, teeth, and postcranial material A rhabdodontid ornithopodR. sp Basségat, La Bastide Neuve, Métisson Upper Multiple specimens StruthiosaurusGarcia, Géraldine; Suberbiola, Xabier Pereda (11 April 2003). "A new species of Struthiosaurus (Dinosauria: Ankylosauria) from the Upper Cretaceous of Villeveyrac (southern France)". Journal of Vertebrate Paleontology. 23 (1): 156–165. doi:10.1671/0272-4634(2003)23[156:ANSOSD]2.0.CO;2. S2CID 140174401. S. languedocensis L'Olivet Quarry, Villeveyrac Upper A partial skeleton consists of a complete synsacrum, pelvic girdle, and four caudal-most dorsal vertebrae A struthiosaurin nodosaurine
Sauropods Sauropods of the Argiles et Grès à Reptiles Formation Genus Species Location Stratigraphic position Material Notes Images Ampelosaurus A. atacis Bellevue locality Upper Three dorsal vertebrae A lirainosaurine lithostrotian titanosaur Atsinganosaurus A. velauciensis La Bastide Neuve, Velaux Lower/Upper A partial skeleton consists of vertebrae, teeth, and sacrum. A lirainosaurine lithostrotian titanosaur Garrigatitan G. meridionalis Velaux-La-Bastide Neuve Upper A holotype consists of a cervical rib, a right humerus, a right ulna, and a left femur. A lirainosaurine lithostrotian titanosaur Hypselosaurus H. priscus Lower A partial femur, fibula and possible tibia and a pair of associated caudal vertebrae. A titanosaurian sauropod
Theropods Theropods of the Argiles et Grès à Reptiles Formation Genus Species Location Stratigraphic position Material Notes Images Abelisauridae indet. Indeterminate A large abelisaurid that was similar to Arcovenator. Arcovenator A. escotae Jas Neuf Sud Lower A complete basicranium articulated with its right postorbital, a tooth, a left squamosal, an anterior caudal vertebra and a right tibia. A large majungasaurine abelisaurid Castignovolucris C. sebei Castigno valley Lower A right coracoid An enantiornithean bird Gargantuavis G. philoinos Lower A fragmentary synsacrum A large avialan Martinavis M. cruzyensis Massecaps locality Lower A complete right humerus An enantiornithean bird Pyroraptor P. olympius La Boucharde locality Lower/Upper Paratypes consist of the equivalent claw of the right foot, the left second metatarsal, a complete second toe claw, a right ulna (long forearm bone), two teeth, five pedal digits, one manual digit, a piece of a metacarpal, a right radius, a dorsal vertebra, and a tail vertebra. A dromaosaurid theropod Theropoda indet. Indeterminate Previously referred to "Megalosaurus" pannoniensis. Variraptor V. mechinorum La Bastide Neuve Lower A posterior dorsal vertebra, a sacrum with five fused vertebrae, and an ilium A dromaeosaurid theropod
Pterosaurs Pterosaurs of the Argiles et Grès à Reptiles Formation Genus Species Location Stratigraphic position Material Notes Images Mistralazhdarcho M. maggii Velaux–La Bastide Neuve Lower A partial skeleton An azhdarchid pterosaur
Crocodilians Crocodilians of the Argiles et Grès à Reptiles Formation Genus Species Location Stratigraphic position Material Notes Images Acynodon A. iberoccitanus A eusuchian crocodylomorph Alligatoridae indet. Indeterminate Allodaposuchus A. iberoarmoricanus Velaux-La Bastide Neuve Lower A partial skull An allodaposuchid eusuchian Crocodylidae indet. Indeterminate Lohuecosuchus L. mechinorum Skull & mandible An allodaposuchid eusuchian
Turtles Turtles of the Argiles et Grès à Reptiles Formation Genus Species Location Stratigraphic position Material Notes Images Calissounemys C. matheroni A compsemydid turtle Foxemys F. mechinorum A bothremydid turtle Polysternon P. provinciale A bothremydid turtle
Fish Fishes of the Argiles et Grès à Reptiles Formation Genus Species Location Stratigraphic position Material Notes Images Hybodontiformes indet. Indeterminate Lepisosteidae indet. Indeterminate
Mammals Mammals of the Argiles et Grès à Reptiles Formation Genus Species Location Stratigraphic position Material Notes Images Mistralestes M. arcensisLa Cairanne-Highway Lower A mandible (Right dentary fragment with p5 to m3 and roots of p4). A zhelestid eutherian Valentinella V. vitrollense Vitrolles-La Plaine Damaged upper and lower jaw fragments. A zhelestid eutherian
Fossil eggs Eggs of the Argiles et Grès à Reptiles Formation Genus Species Location Stratigraphic position Material Notes Images Megaloolithus M. aureliensis M. petralta M. sp.
Arthropods Arthropods of the Argiles et Grès à Reptiles Formation Genus Species Location Stratigraphic position Material Notes Images Dinocarcinus D. velauciensis Velaux-La Bastide Neuve Isolated claws A large-sized portunoid crab
Plants Plants of the Alcântara Formation Genus Species Location Stratigraphic position Material Notes Images Amblyochara A. begudiana Peckichara P. cancellata P. pectinata Platychara P. caudata Saportanella S. maslovi
See also List of dinosaur-bearing stratigraphic units List of fossiliferous stratigraphic units in France
References
Bibliography
Further reading
R. Cousin, G. Breton, R. Fournier and J.-P. Watte. 1994. Dinosaur egglaying and nesting in France. K. Carpenter, K.F. Hirsch & J.R. Horner (eds.) Dinosaur Eggs and Babies. Cambridge University Press, Cambridge 56–74 J. Le Loeuff, E. Buffetaut, P. Méchin and A. Méchin-Salessy. 1992. The first record of dromaeosaurid dinosaurs (Saurischia, Theropoda) in the Maastrichtian of southern Europe: palaeobiogeographical implications. Bulletin de la Société géologique de la France 163(3):337-343 R. Dughi and F. Sirugue. 1958. Sur les oeufs de dinosaures du bassin fluvio-lacustre de Basse-Provence. Comptes Rendus du Congrès National des Sociétés Savantes, Section des Sciences, Paris 83:183-20