Arachniodes is a fern genus in the family Dryopteridaceae (wood ferns), subfamily Dryopteridoideae, in the Pteridophyte Phylogeny Group classification of 2016 (PPG I). A number of species in this genus are known as "holly ferns".Arachniodes Flora of North America 26 Jan 2012
The genus Arachniodes was first published by Carl Ludwig von Blume in 1828, with the single Indonesian species Arachniodes aspidioides.Blume, Carl (Karl) Ludwig von; Enumeratio Plantarum Javae fasc. 2: 241. 1828. The genus was not widely recognized until Mary Douglas Tindale transferred the two species (Byrsopteris amabilis and Byrsopteris aristata) into it in 1961.Tindale, Mary Douglas; Contributions from the New South Wales National Herbarium 3(1): 90. 1961.
Species The Pteridophyte Phylogeny Group classification of 2016 (PPG I) considers Arachniodes to have 60 species. , the Checklist of Ferns and Lycophytes of the World accepted 78 species:
Arachniodes ailaoshanensis Ching Arachniodes amabilis (Blume) Tindale Arachniodes amoena (Ching) Ching Arachniodes aristata (G.Forst.) Tindale Arachniodes assamica (Kuhn) Ohwi Arachniodes bella (C.Chr.) Ching Arachniodes blinii (H.Lév.) Nakaike Arachniodes carvifolia (Kunze) Ching Arachniodes caudata Ching Arachniodes cavaleriei (Christ) Ohwi Arachniodes chinensis (Rosenst.) Ching Arachniodes coniifolia (T.Moore) Ching Arachniodes cornucervi (D.Don) Fraser-Jenk. Arachniodes daklakensis Li Bing Zhang, N.T.Lu & X.M.Zhou Arachniodes davalliaeformis (Christ) Nakaike Arachniodes denticulata (Sw.) Ching Arachniodes fengii Ching Arachniodes festina (Hance) Ching Arachniodes formosa (Fée) Ching Arachniodes formosissima (Goldm.) Nakaike Arachniodes gigantea Ching Arachniodes globisora (Hayata) Ching Arachniodes grossa (Tardieu & C.Chr.) Ching Arachniodes hainanensis (Ching) Ching Arachniodes haniffii (Holttum) Ching Arachniodes hehaii Li Bing Zhang, N.T.Lu & X.F.Gao Arachniodes hekiana Kurata Arachniodes henryi (Christ) Ching Arachniodes hiugana Sa. Kurata Arachniodes hunanensis Ching Arachniodes insularis W.H.Wagner Arachniodes japonica (Kurata) Nakaike Arachniodes jinpingensis Y.T.Hsieh Arachniodes leucostegioides (C.Chr.) Ching Arachniodes longicaudata Li Bing Zhang, N.T.Lu & Liang Zhang Arachniodes longipinna Ching Arachniodes microlepioides (C.Chr.) comb. ined. Arachniodes miqueliana (Maxim. ex Franch. & Sav.) Ohwi Arachniodes miyakei (H.Itô) Shimura ex Nakaike Arachniodes mutica (Franch. & Sav.) Ohwi Arachniodes neopodophylla (Ching) Nakaike Arachniodes nigrospinosa (Ching) Ching Arachniodes nipponica (Rosenst.) Ohwi Arachniodes oosorae H.Itô Arachniodes palmipes (Kunze) Fraser-Jenk. Arachniodes pseudoaristata (Tagawa) Ohwi Arachniodes pseudoassamica Ching Arachniodes pseudorepens Nakaike Arachniodes puncticulata (Alderw.) Ching Arachniodes quadripinnata (Hayata) Seriz. Arachniodes quangnamensis Li Bing Zhang, N.T.Lu & Liang Zhang Arachniodes repens Kurata Arachniodes rhomboidea (Schott) Ching Arachniodes rigidissima (Hook.) Ching Arachniodes sarasiniorum (Christ) Nakaike Arachniodes simplicior (Makino) Ohwi Arachniodes simulans (Ching) Ching Arachniodes sinomiqueliana (Ching) Ohwi Arachniodes sinorhomboidea Ching Arachniodes sledgei Fraser-Jenk. Arachniodes spectabilis (Ching) Ching Arachniodes squamulosa R.C.Moran & B. Øllg. Arachniodes standishii (T.Moore) Ohwi Arachniodes subamabilis Kurata Arachniodes superba Fraser-Jenk. Arachniodes tomitae Sa. Kurata Arachniodes tonkinensis (Ching) Ching Arachniodes tripinnata (Goldm.) Sledge Arachniodes tsiangiana (Ching) Nakaike Arachniodes webbiana (A.Braun) Schelpe Arachniodes wulingshanensis S.F.Wu Arachniodes yakusimensis (H.Itô) Nakaike Arachniodes ziyunshanensis Y.T.Xie
Some hybrids were also accepted:
Arachniodes × azuminoensis Fujiw., Y.Matsuda, Yu. Abe & Otsuka Arachniodes × chibaensis Yashiro Arachniodes × clivorum Kurata Arachniodes × ikeminensis Seriz. Arachniodes × kenzo-satakei (Kurata) Kurata Arachniodes × kurosawae Shimura & Kurata Arachniodes × masakii Kurata Arachniodes × minamitanii Sa. Kurata Arachniodes × mirabilis Sa. Kurata Arachniodes × mitsuyoshiana Sa. Kurata Arachniodes × pseudohekiana Kurata Arachniodes × respiciens Sa. Kurata Arachniodes × sahashii Fraser-Jenk. & Kandel Arachniodes × sasamotoi Kurata Arachniodes × takayamensis Seriz.
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