Bythinella is a genus of very small freshwater snails, an aquatic gastropod mollusk in the family Bythinellidae (according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005).
These gonochoristic snails are small animals, with a shell length of about 2–4 mm.Bichain et al. (2007) Amnicola is a close relative, containing North American snails with essentially similar appearance and ecology. The scientific name means "small Bithynia"; the latter snail genus is somewhat similar at a casual glance (though much larger) but not at all closely related among the Rissooidea.
Distribution The present genus occurs in springs, and sometimes caves or groundwater, from Catalonia across central Europe to the Aegean Region of Turkey. In and around France a particularly high diversity of these snails is found. Due to the limited range and special and easily destroyed habitat, these snails are liable to become endangered species.
Species More than 80 species and subspecies are recognized at present. However, it is not at all clear that all of those recognized as good species are actually that distinct, and some in fact seem to be local morphs instead of valid taxa. Some undescribed species are also known to exist.
Bythinella abbreviata Michaud, 1831 Bythinella alexpeteri Glöer & Hirschfelder, 2020 †Bythinella alta Deshayes, 1862 Bythinella amira Glöer & Hirschfelder, 2020 Bythinella anatolica Yıldırım, Kebapçı & Bahadır Koca, 2015 Bythinella andorrensis Paladilhe, 1875 Bythinella aneliae Georgiev & Stoycheva, 2011 Bythinella angelitae Haase, Wilke & Mildner, 2007 Bythinella angelovi Glöer & Georgiev, 2011 Bythinella angusta Clessin, 1911 Bythinella anianensis Paladilhe, 1870 - accepted as: B. cebennensis † Bythinella antiguensis A. P. Brown & Pilsbry, 1914 † Bythinella archaea Bourguignat, 1869 † Bythinella atomus Brongniart, 1810 Bythinella atypikos A. Reischütz, P. L. Reischütz & W. Fischer, 2008 † Bythinella auriculata Szőts, 1953 Bythinella austriaca Frauenfeld, 1856 Bythinella badensis Boeters, 1981: accepted as Bythinella reyniesii Bythinella baidashnikovi Sitnikova, Starobogatov & V. Anistratenko, 1992 Bythinella batalleri Bofill, 1925 Bythinella baudoni (Paladilhe, 1874) Bythinella bavarica Clessin, 1877 † Bythinella bearnensis Cossmann & Peyrot, 1918 Bythinella beckmanni A. Reischütz, P. L. Reischütz & W. Fischer, 2008 Bythinella bertrandi R. Bernasconi, 2000 Bythinella bicarinata (Des Moulins, 1827) Bythinella blidariensis Glöer, 2013 Bythinella bouldouyrensis Girardi, 2015 Bythinella bouloti Girardi, Bichain & Wienin, 2002 Girardi H., Bichain J. M. & Wienin M. (2002) "Description de deux nouvelles espèces de Bythinella (Mollusca: Caenogastropoda: Hydrobiidae) de l’exsurgence de Castelbouc (Lozère, France)". Documents Malacologiques 3: 43-54. Bythinella calimanica Falniowski, Szarowska & Sirbu, 2009 Bythinella calquierensis Girardi & Boeters, 2015 Bythinella carcasonis Boeters & Falkner, 2008 Bythinella carinulata (Drouët, 1867): accepted as Bythinella viridis (Poiret, 1801) Bythinella cebennensis (Dupuy, 1851) Bythinella charpentieri (Roth, 1855) † Bythinella cirsophora Cossmann, 1888 Bythinella collingi Boeters, 2009 Bythinella companyoi (Paladilhe, 1870) Bythinella conica Clessin, 1910 † Bythinella contemta Brusina, 1897 Bythinella cretensis Schütt, 1980 Bythinella curta Clessin, 1911 † Bythinella cyclothyra (Boettger, 1869) Bythinella cylindracea (Paladilhe, 1869) Bythinella cylindrica (Frauenfeld, 1857) Bythinella dacica Grossu, 1946 † Bythinella dalmatica Brusina, 1897 † Bythinella damintunensis Youluo, 1978 Bythinella darrieuxii (de Folin & Bérillon, 1877) Bythinella dedovi Glöer & Georgiev, 2011 Bythinella deidamiae Glöer & Porfyris, 2022 † Bythinella deshayesiana (Bourguignat, 1869) † Bythinella desnoyersi (Bourguignat, 1869) Bythinella dierckingi Glöer & Georgiev, 2011 Bythinella dispersa Radoman, 1976 Bythinella drimica Radoman, 1976 Bythinella dromensis Boeters & Falkner, 2008 † Bythinella dumenisliana (Bourguignat, 1869) † Bythinella edwardsiana (Bourguignat, 1869) Bythinella ehrmanni Pax, 1938 Bythinella elenae Glöer & Georgiev, 2011 Bythinella eleousae Glöer & Hirschfelder, 2020 Bythinella ellinikae Glöer & Hirschfelder, 2020 † Bythinella elongata Roshka, 1973 Bythinella espanoli Bech, 1980 Bythinella etrusca (Paladilhe, 1867) † Bythinella eugenii (Deshayes, 1862) Bythinella eurystoma (Paladilhe, 1870) Bythinella eutrepha (Paladilhe, 1867) † Bythinella expulsa (Deshayes, 1862) † Bythinella eysdenensis É. Vincent, 1930 Bythinella falniowskii Glöer, 2013 Bythinella feheri Glöer, 2013 Bythinella ferussina (Des Moulins, 1827) Bythinella friderici Boeters & Falkner, 2008 Bythinella fuscata Brancsik, 1889 Bythinella galerae Girardi, Bichain & Wienin, 2002: accepted as Bythinella bouloti H. Girardi, Bichain & Wienin, 2002 Bythinella geisserti Boeters & Falkner, 2003 Bythinella georgievi Glöer, 2013 Bythinella gibbosa (Moquin-Tandon, 1856) Bythinella gloeeri Georgiev, 2009 Bythinella gokceadaensis D. A. Odabaşı, 2019 Bythinella golemoensis Glöer & Mrkvicka, 2015 † Bythinella gracillima Szőts, 1953 Bythinella gregoi Glöer & Erőss, 2015 Bythinella grossui Falniowski, Szarowska & Sirbu, 2009 Bythinella guranensis (Paladilhe, 1870): accepted as Bythinella rubiginosa (Boubée, 1833) † Bythinella hanjiangensis W. Yü, 1977 Bythinella hansboetersi Glöer & Pešić, 2006 Bythinella hasanspahici Mulaomerović & Glöer, 2022 Bythinella heydeni Clessin, 1879 Bythinella heynemanniana Hazay, 1881 Bythinella hungarica Hazay, 1880 † Bythinella intermedia (Melleville, 1843) Bythinella istanbulensis Yıldırım, Kebapçı & Yüce, 2015 Bythinella istoka Glöer & Pešić, 2014 Bythinella izvorica Glöer & Georgiev, 2011 Bythinella jobae (Bourguignat, 1862) Bythinella jodevidtsi Bichain & Bertrand, 2022 Bythinella joinvillensis (Bourguignat, 1869) Bythinella jourdei R. Bernasconi, 2000 Bythinella jozefgregoi Glöer & Reuselaars, 2020 Bythinella kambosensis Glöer & Hirschfelder, 2020 Bythinella kapelana Radoman, 1976 Bythinella kastaliae Glöer & Hirschfelder, 2020 Bythinella kastanolongosensis Glöer & Pešić, 2020 Bythinella kazdaghensis Odabaşi & Georgiev, 2014 Bythinella kleptuzica Glöer & Georgiev, 2011 Bythinella klimaensis Glöer & Reuselaars, 2020 Bythinella konstadinensis Glöer & Reuselaars, 2020 Bythinella kosensis Schütt, 1980 Bythinella kosmosi Clessin, 1910 Bythinella kwanti Glöer & Reuselaars, 2020 Bythinella kyriaki Glöer & Reuselaars, 2020 Bythinella lancevelei Locard, 1894 accepted as Bythinella viridis (Poiret, 1801) †Bythinella lartetiana (Bourguignat, 1869) †Bythinella lecongensis W. Yü & X.-Q. Zhang, 1982 Bythinella liandinaensis Glöer & Reuselaars, 2020 Bythinella ligurica (Paladilhe, 1867) Bythinella limnopsis Letourneux & Bourguignat, 1887 Bythinella longula Brancsik, 1889 Bythinella lunzensis Boeters, 2008 Bythinella luteola Radoman, 1976 Bythinella magdalenae Yıldırım, Kebapçı & Bahadır Koca, 2015 Bythinella magna Radoman, 1976 Bythinella major (Pascal, 1873) Bythinella margritae Glöer & Georgiev, 2011 Bythinella marianramosae Falniowski, Jaszczyńska & Hofman, 2023 Bythinella marici Glöer & Pešić, 2014 Bythinella markovi Glöer & Georgiev, 2009 Bythinella mauritanica Letourneux & Bourguignat, 1887 † Bythinella megarensis Bukowski, 1896 Bythinella melovskii Glöer & Slavevska-Stamenković, 2015 Bythinella metarubra Falniowski, 1987 Bythinella micherdzinskii Falniowski, 1980 Bythinella microcochlia Letourneux & Bourguignat, 1887 †Bythinella minuata (Deshayes, 1862) Bythinella molcsanyi H. Wagner, 1941 Bythinella moulinsii (Dupuy, 1849) Bythinella muranyii Glöer & Erőss, 2015 Bythinella navacellensis Prié & Bichain, 2009 Bythinella nonveilleri Glöer, 2008 Bythinella normalis Courty, 1907 Bythinella nothites (Westerlund, 1886) Bythinella occasiuncula Boeters & Falkner, 2001 Bythinella olymbosensis Glöer & Reuselaars, 2020 Bythinella opaca (M. von Gallenstein, 1848) Bythinella padana R. Bernasconi, 1989 Bythinella padiraci Locard, 1903 Bythinella pannonica (Frauenfeld, 1865) †Bythinella parkinsoni (J. Morris, 1854) Bythinella parvula Locard, 1893: accepted as Bythinella reyniesii (Dupuy, 1851) Bythinella perilongata Altimira, 1959 Bythinella perivoliensis Glöer & Reuselaars, 2020 Bythinella pesici Glöer & Reuselaars, 2020 Bythinella pesterica Glöer, 2008 Bythinella petrosensis Glöer, 2008 Bythinella pujolensis R. Bernasconi, 2000: accepted as Bythinella bicarinata (Des Moulins, 1827) †Bythinella pulcherrima Szőts, 1953 †Bythinella pulchra (Deshayes, 1862) Bythinella punica Letourneux & Bourguignat, 1887 †Bythinella pupina (Deshayes, 1862) Bythinella pupoides (Paladilhe, 1869) Bythinella pyrenaica (Bourguignat, 1861) Bythinella rachonica Georgiev & Glöer, 2020 Bythinella radomani Falniowski, Szarowska & Sirbu, 2009 Bythinella ravnogorica Glöer & Georgiev, 2009 Bythinella reischuetzi Glöer & Georgiev, 2020 Bythinella rethymnonensis Glöer & Hirschfelder, 2020 Bythinella reuselaarsi Glöer & Pešić, 2020 Bythinella reyniesii (Dupuy, 1851) †Bythinella rhodanica Roman, 1912 Bythinella rhodopensis Glöer & Georgiev, 2011 Bythinella righaensis Glöer & Reuselaars, 2020 Bythinella rilaensis Georgiev & Glöer, 2013 †Bythinella rimulifera Stache, 1889 Bythinella robiciana Bythinella robusta Z.-B. Kang, 1986 Bythinella rolani Boeters, 2019 Bythinella rondelaudi R. Bernasconi, 2000 † Bythinella rothi (Brusina, 1884) Bythinella roubionensis Caziot, 1910 Bythinella rouchi Boeters & Falkner, 2008 Bythinella rouveyrolensis Girardi, 2015 Bythinella rubiginosa (Boubee, 1833) Bythinella reyniesii (Dupuy, 1851) Bythinella samecana Clessin, 1911 †Bythinella scalaris (Slavík, 1869) Bythinella schmidtii (Küster, 1852): accepted as Bythinella opaca (M. von Gallenstein, 1848) † Bythinella scitula Brusina, 1892 Bythinella serborientalis Radoman, 1976 Bythinella servainiana (Paladilhe, 1870) Bythinella sijei Mulaomerović & Glöer, 2022 †Bythinella sinensis W. Yü & H.-Z. Pan, 1982 Bythinella sirbui Glöer, 2013 Bythinella sitiensis Glöer & Georgiev, 2011 Bythinella slaveyae Glöer & Georgiev, 2011 Bythinella smolyanica Glöer & Georgiev, 2011 Bythinella solidula Brancsik, 1889 Bythinella sordida (Küster, 1853) †Bythinella sphaeroidalis Cossmann, 1888 Bythinella srednogorica Glöer & Georgiev, 2009 Bythinella steffeki Grego & Glöer, 2019 Bythinella stoychevae Georgiev, 2011 Bythinella subovata (Paladilhe, 1876) Bythinella syngenes Preston, 1914 Bythinella syntriculus Boeters & Falkner, 2008 Bythinella szarowskae Glöer, 2013 Bythinella tabakovaensis Mulaomerović & Glöer, 2023 Bythinella tajoensis Boeters, 2019 Bythinella taraensis Glöer & Pešić, 2010 Bythinella taygetensis Glöer & Hirschfelder, 2020 Bythinella tejedoi Boeters, 2019 Bythinella temelkovi Georgiev & Glöer, 2014 Bythinella thermophila Glöer, Varga & Mrkvicka, 2015 Bythinella tornensis Hazay, 1881 Bythinella troyana R. Bernasconi, 2000 Bythinella truncata Courty, 1907 Bythinella tumidula Clessin, 1910 Bythinella turca (Radoman, 1976) Bythinella turriculata (Paladilhe, 1869) Bythinella ullaensis Boeters & Falkner, 2008 Bythinella utriculus (Paladilhe, 1874) Bythinella valkanovi Glöer & Georgiev, 2011 Bythinella vesontiana R. Bernasconi, 1989 Bythinella vidinovae Dedov, Taseva & Georgiev, 2021 Bythinella vimperei R. Bernasconi, 2000 Bythinella viridis (Poiret, 1801) Bythinella viseuiana Falniowski, Szarowska & Sirbu, 2009 † Bythinella vitrellaeformis Lörenthey, 1902 Bythinella walensae Falniowski, Hofman & Rysiewska, 2016 Bythinella walkeri Glöer & Georgiev, 2009 Bythinella wawrzineki R. Bernasconi, 2002 Bythinella westerlundi Glöer, 2022 Bythinella wilkei Yıldırım, Kebapçı & Bahadır Koca, 2015 Bythinella yerlii Gürlek, 2017 Bythinella zyvionteki Falniowski, 1987
Relocated species: