Helicoidea is a taxonomic superfamily of air-breathing land snails, terrestrial pulmonate gastropod mollusks in the order Stylommatophora.
Taxonomy
2017 taxonomy and latest developments Sources:
The Bouchet et al. 2017 nomenclator provides an up to date system of Helicoidea. The system is in some parts preliminary, as the authors relied on unpublished (as of 2023) phylogenomic study, which did not include all New World taxa. They classified Epiphragmophoridae, Helminthoglyptidae, Humboldtianidae, Monadeniidae and Xanthonychidae (provisionally also Lysinoidae and Echinichidae) as subfamilies of the last taxon, because there are no deep splits between them in the cited unpublished study (see also Koene & Schulenburg 2005).
Cepolidae, Labyrinthidae and Thysanophoridae constitute the sister group of the remaining Helicoidea. Camaenidae Pilsbry, 1895 Canariellidae Schileyko, 1991 Cepolidae Ihering, 1909 Elonidae Gittenberger, 1977 Geomitridae C. Boettger, 1909 Helicidae Rafinesque, 1815 Helicodontidae Kobelt, 1904 Hygromiidae Tryon, 1866 Labyrinthidae Borrero, Sei, Robinson & Rosenberg, 2017 Genus: † Laxecostula Harzhauser & Neubauer, 2020 Pleurodontidae Ihering, 1912 Polygyridae Pilsbry, 1895 Sphincterochilidae Zilch, 1960 Thysanophoridae Pilsbry, 1926 Trissexodontidae H. Nordsieck, 1987 Xanthonychidae Strebel & Pfeffer, 1879 Trichodiscinidae H. Nordsieck, 1987
2015 taxonomy Razkin et al. (2015)Razkin O., Gómez-Moliner B. J., Prieto C. E., Martínez-Ortí A., Arrébola J. R., Chueca L. J. & Madeira M. J. (2015). "Molecular phylogeny of the western Palearctic Helicoidea (Gastropoda, Stylommatophora)". Molecular Phylogenetics and Evolution 83: 99–117. . reorganized classification into monophyletic taxa according to the molecular phylogeny. This study is focus on Western Palaearctic species. The Hygromiidae s.l. family was divided into three families, Canariellidae, Geomitridae and Hygromiidae. Moreover, the family Cochlicellidae was including within the Geomitridae family as a tribe (Cochlicellini).
The classification proposed for Western Palaearctic Helicoidea is as follows: Bradybaenidae (now Bradybaeninae - subfamily of Camaenidae) Camaenidae Canariellidae Elonidae Geomitridae Geomitrinae Cochlicellini Geomitrini Plentuisini Ponentinini Helicellinae Cernuellini Helicellini Trochoideini Helicodontidae Helicidae Ariantinae Helicinae Allognathini Helicini Otalini Thebini Murellinae Humboldtianidae Hygromiidae Ciliellinae Hygromiinae Leptaxinae Sphincterochilidae Trissexodontidae Gittenbergeriinae Trissexodontinae
2012 taxonomy Thompson & Naranjo-García (2012) described a new family Echinichidae and placed it to the superfamily Xanthonychoidea.Thompson F. G. & Naranjo-García E. (2012). "Echinichidae, a new family of dart-bearing helicoid slugs from Mexico, with the description of a new genus and three new species (Gastropoda: Pulmonata: Xanthonychoidea)". Archiv für Molluskenkunde 141(2): 197–208. . preview. Therefore, the family Xanthonychidae was moved from Helicoidea to Xanthonychoidea.
2005 taxonomy There are 19 families within the superfamily Helicoidea according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005.Poppe G. T. & Tagaro S. P. (2006). "The new classification of Gastropods according to Bouchet & Rocroi, 2005". Visaya, février 2006: 10 pp. PDF . Helicidae Bradybaenidae Camaenidae Cepolidae Cochlicellidae Elonidae Epiphragmophoridae Halolimnohelicidae Helicodontidae Helminthoglyptidae Humboldtianidae Hygromiidae Monadeniidae Pleurodontidae Polygyridae Sphincterochilidae Thysanophoridae Trissexodontidae Xanthonychidae
According to H. NordsieckHigher classification of the Helicoidea and the molecular analyses of their phylogeny (by Hartmut Nordsieck) the family Xanthonychidae (sensu Hausdorf & Bouchet) is probably polyphyletic (contains several different lineages) and therefore should be divided into several families.
References