The California condor (Gymnogyps californianus) is a New World vulture and the largest North American land bird. It became extinct in the wild in 1987 when all remaining wild individuals were captured, but has since been reintroduced to northern Arizona and southern Utah (including the Grand Canyon area and Zion National Park), the coastal mountains of California, and northern Baja California in Mexico. It is the only surviving member of the genus Gymnogyps, although four extinct members of the genus are also known. The species is listed by IUCN as critically endangered, and similarly considered Critically Imperiled by NatureServe.
The plumage is black with patches of white on the underside of the wings; the head is largely bald, with skin color ranging from gray on young birds to yellow and bright orange on breeding adults. Its wingspan is the widest of any North American bird, and its weight of up to nearly equals that of the trumpeter swan, the heaviest among native North American bird species. The condor is a scavenger and eats large amounts of carrion. It is one of the world's longest-living birds, with a lifespan of up to 60 years.
Condor numbers dramatically declined in the 20th century due to agricultural chemicals (DDT), poaching, lead poisoning, and habitat destruction. A conservation plan put in place by the United States government led to the capture of all the remaining wild condors by 1987, with a total population of 27 individuals. These surviving birds were bred at the San Diego Wild Animal Park and the Los Angeles Zoo. Numbers rose through captive breeding, and beginning in 1991, condors were reintroduced into the wild. Since then, their population has grown, but the California condor remains one of the world's rarest bird species. In December 2025, the Fish and Wildlife Service updated the world population to 607. The condor is a significant bird to many Californian Native American groups and plays an important role in several of their traditional myths.
Taxonomy
The California condor was described by English naturalist George Shaw in 1797 as Vultur californianus; Archibald Menzies collected the type specimen "from the coast of California" during the Vancouver Expedition. It was originally classified in the same genus as the Andean condor (V. gryphus), but, due to the Andean condor's slightly different markings, slightly longer wings, and tendency to kill small animals to eat,Nielsen 2006, p. 27 the California condor has been placed in its own monotypic genus. The generic name Gymnogyps is derived from the Greek gymnos/γυμνος "naked" or "bare", and gyps/γυψ "vulture", while the specific name californianus comes from its location in California. The word condor itself is derived from the Quechua word kuntur.
The exact taxonomic placement of the California condor and the other six species of New World vultures remains unclear. Though similar in appearance and ecological roles to Old World vultures, the New World vultures evolved from a different ancestor in a different part of the world. Just how different the two are is under debate, with some earlier authorities suggesting that the New World vultures are more closely related to storks.Sibley, Charles G. and Monroe, Burt L. 1990. Distribution and Taxonomy of the Birds of the World . Yale University Press. . Accessed April 11, 2007. More recent authorities maintain their overall position in the order Falconiformes along with the Old World vulturesSibley, Charles G., and Ahlquist, Jon E. 1991. Phylogeny and Classification of Birds: A Study in Molecular Evolution . Yale University Press. . Accessed April 11, 2007. or place them in their own order, Cathartiformes. Electronic Supplementary Material (PDF) The South American Classification Committee has removed the New World vultures from Ciconiiformes and instead placed them in Incertae sedis, but notes that a move to Falconiformes or Cathartiformes is possible.Remsen, J. V. Jr.; C. D. Cadena; A. Jaramillo; M. Nores; J. F. Pacheco; M. B. Robbins; T. S. Schulenberg; F. G. Stiles; D. F. Stotz & K. J. Zimmer. 2007. A Classification of the Bird Species of South America. South American Classification Committee. Retrieved on October 15, 2007
As of the 51st Supplement (2010) of the American Ornithologists' Union, the California condor is in the family Cathartidae of the order Cathartiformes.
Evolutionary history
The genus Gymnogyps is an example of a relict distribution. During the Pleistocene Epoch, this genus was widespread across the Americas. From fossils, the Floridian Gymnogyps kofordi from the Early Pleistocene and the Peruvian Gymnogyps howardae from the Late Pleistocene have been described. A condor found in Late Pleistocene deposits on Cuba was initially described as Antillovultur varonai, but has since been recognized as another member of Gymnogyps, Gymnogyps varonai. It may even have derived from a founder population of California condors.
The California condor is the sole surviving member of Gymnogyps and has no accepted subspecies. However, there is a Late Pleistocene form that is sometimes regarded as a palaeosubspecies, Gymnogyps californianus amplus. Opinions are mixed, regarding the classification of the form as either a chronospecies or a separate species, Gymnogyps amplus. Gymnogyps amplus occurred over much of the bird's historical range – even extending into Florida – but was larger, having about the same weight as the Andean condor. This bird also had a wider bill. As the climate changed during the last ice age, the entire population may have become smaller until it had evolved into the Gymnogyps californianus of today, although more recent studies by Syverson question that theory.
Description
The adult California condor is a uniform black with the exception of large triangular patches or bands of white on the underside of the wings. It has gray legs and feet, an ivory-colored bill, a frill of black feathers surrounding the base of the neck, and brownish red eyes. The juvenile is mostly a mottled dark brown with blackish coloration on the head. It has mottled gray instead of white on the underside of its flight feathers.BirdLife International (2007) Species factsheet: California Condor Gymnogyps californianus . Retrieved August 17, 2007
The condor's head has little to no feathers, which helps keep it clean when feeding on carrion. The skin of the head and neck is capable of flushing noticeably in response to emotional state. The skin color varies from yellowish to a glowing reddish-orange. The birds do not have true syringeal vocalizations. They can make a few hissing or grunting sounds only heard when very close.
The female condor is smaller than the male, an exception to the rule among birds of prey (the related Andean condor is another exception). Overall length ranges from and wingspan from . Their weight ranges from , with estimations of average weight ranging from .Raptors of the World by Ferguson-Lees, Christie, Franklin, Mead & Burton. Houghton Mifflin (2001), Wingspans of up to have been reported but no wingspan over has been verified. Most measurements are from birds raised in captivity, so it is difficult to determine if major differences exist between wild and captive condors.
California condors have the largest wingspan of any North American bird. They are surpassed in both body length and weight only by the trumpeter swan and the introduced mute swan. The American white pelican and whooping crane also have longer bodies than the condor. Condors are so large that they can be mistaken for a small, distant airplane, which possibly occurs more often than that they are mistaken for other bird species.Nielsen 2006, p. 1
The middle toe of the California condor's foot is greatly elongated, and the hind toe is only slightly developed. The talons of all the toes are straight and blunt and are thus more adapted to walking than gripping. This is more similar to their supposed relatives the storksCracraft, J. et al. 2004. "Phylogenetic relationships among modern birds (Neornithes): toward an avian tree of life". pp. 468–489 in Assembling the Tree of Life (Cracraft, J. and Donoghue, M. J. eds.). Oxford University Press, New York. . than to birds of prey and Old World vultures, which use their feet as weapons or organs of prehension.
Historic range
At the time of human settlement of the Americas, the California condor was widespread across North America; condor bones from the late Pleistocene have been found at the Cutler Fossil Site in southern Florida. However, at the end of the last glacial period came the extinction of the megafauna that led to a subsequent reduction in range and population. Five hundred years ago, the California condor roamed across the American Southwest and West Coast. Faunal remains of condors have been found documented in Arizona, Nevada, New Mexico, and Texas.
The Lewis and Clark Expedition of the early 19th century reported on their sighting and shooting of California condors near the mouth of the Columbia River. The species was observed as far north as Lulu Island, British Columbia in the late 1880s; it was described as once being "common" in the area up until that point. Its 19th century presence at the mouth of the Fraser River was corroborated by John Keast Lord.
In the 1970s, two Condor Observation Sites were established in the Santa Clara River Valley to host hopeful birders interested in the endangered species: one about 15 miles north of Fillmore, California, near the Sespe Wildlife Area of Los Padres National Forest, and one atop Mount Pinos, "accessible from a dirt road off the highway in from Gorman".
Habitat The California condor lives in rocky shrubland, coniferous forest, and oak savanna. They are often found near cliffs or large trees, which they use as nesting sites. Individual birds have a huge range and have been known to travel up to in search of carrion.
There are two sanctuaries chosen because of their prime condor nesting habitat: the Sisquoc Condor Sanctuary in the San Rafael WildernessGagnon, Dennis R. Hiking the Santa Barbara Backcountry. The Ward Ritchie Press, Pasadena, California, 1974. and the Sespe Condor Sanctuary in the Los Padres National Forest.
The Los Padres Condor Range and River Protection Act of 1992 expanded existing wilderness by and designated of new wilderness that provide habitat for the condor in the Los Padres.
Ecology and behavior
The California condor's large flight muscles are not anchored by a correspondingly large sternum, which restricts them to being primarily soarers. The birds flap their wings when taking off from the ground, but after attaining a moderate elevation they largely glide, sometimes going for miles without a single flap of their wings. They have been known to fly up to speeds of and as high as . They prefer to roost on high perches from which they can launch without any major wing-flapping effort. Often, these birds are seen soaring near rock cliffs, using thermals to aid them in keeping aloft.Nielsen 2006, p. 79
The California condor has a long life span, reaching up to 60 years. If it survives to adulthood, the condor has few natural threats other than humans. Because they lack a syrinx, their vocal display is limited to grunts and hisses. Condors bathe frequently and can spend hours a day preening their feathers. Condors also perform urohidrosis, defecation on their legs, to reduce their body temperature. There is a well-developed social structure within large groups of condors, with competition to determine a pecking order decided by body language, competitive play behavior, and a variety of hisses and grunts. This social hierarchy is displayed especially when the birds feed, with the dominant birds eating before the younger ones.
Breeding